The American Seven-spined Gobies
Tribe Gobiosomatini

Species Documentation


Prepared by:

James Van Tassell

Research Associate in Residence

Dept. of Biology

114 Hofstra University

Hempstead, NY 11549


Revision Date: Aug. 29, 2002

This is a working document and not a completed list of all references, synonoyms, etc.  If you see any mistakes or wish to contribute additional information, please e-mail me.


The following genera are included in alphabetical order:  Aboma,  Akko, Aruma, Barbulifer, Bollmannia , Chriolepis, Eleotrica , Enypnias, Evermannichthys, Ginsburgellus, Gobiosoma (including the subgenera – Austrogobius, Garmannia , Gobiolepis, Gobiosoma, Elacatinus, Tigrigobius ), Gobulus, Gymneleotris, Microgobius, Nes, Ophiogobius, Palatogobius, Pariah, Parrella, Psilotris, Pycnomma, Robinsichtys, Risor, Varicus.



KEY: Information within the brackets [ ] indicates the source of the information and that the information has been checked against the original publication.


NOTE:  The classification used for Gobiosoma is that of  Böhlke & Robins 1968, when ever possible.  While both Hoese (1971) and Van Tassell (1998) have proposed new classifications for all or part of the group in their Ph.D. dissertations, neither is currently published.  When I have sufficient information to resolve a few remaining nodes, I’ll publish a new phylogeny and changes will then be made to this page.


Genus Aboma


Aboma Jordan and Starks 1895; Type species Aboma etheostoma Jordan and Starkes; Fishes of Sinaola, Proc. Calif. Acad. Sci 2d ser., 1895: 497. [Jordan, 1930]


etheostoma Jordan and Starks 1895; locality: Mazatlan, Mexico.

polyporosum Dawson 1969: 510-513; holotype USNM 203573; type locality: Panama, Pacific coast from deep tidepool at Farfan Point.


Genus Akko


Akko Birdsong and Robbins 1995


dionaea Birdsong & Robins 1995: 676, by original designation; holotype USNM 329524; type locality: Amapa Prov. East of Iiha de Maraca, Brazil.


Genus Aruma


Aruma Ginsburg 1933a: 16 Type species: Gobiosoma occidentale Ginsburg 1933:16-17, by original designation; erected as a subgenus of Gobiosoma.[Gin. 1933a]


histrio (Jordan 1884)

Gobiosoma histrio Jordan 1884:260; holotype USNM 35147; Type locality: Guaymas, Mexico [Hoese 1976]{Proc. U.S. Nat. Mus. for 1884 was published in 1885}

Gobiosoma occidentale Ginsburg 1933a; Type USNM 92801; Type locality: La Paz Harbor, Mexico;[Gin. 1933] [syn. by Hoese 1976 Copeia No. 2]


Genus Barbulifer


Barbulifer Eigenmann and Eigenmann 1888(1889): 70 Type species: Barbulifer papillosus Eigenmenn and Eigenmann 1888(1889): 70 [= Gobiosoma ceuthoecum Jordan and Gilbert 1884] by original designation, see Böhlke & Robins 1968: 120 for additional information on problems associated with the type species. [B&R 1968] [Eigenmann and Eigenmann 1888(1889)][Hoese 1985]


ceuthoecus (Jordan and Gilbert 1884)


Gobiosoma ceuthoecum Jordan and Gilbert 1884: 29, holotype USNM 34970; Type locality: Key West, Florida, from the cavity of a sponge. Jordan 1887: 894 [106] (in check=list). Jordan and Eigenmann 1887: 506, 508 (synopsis of species).[B&R 1968] [Jordan and Gilbert 1884]

Barbulifer ceuthoecus: Eigenmann and Eiganmann 1888: 72 (placed in Barbulifer, with papillosus). Jordan and Evermann 1896: 461 (in check-list). Jordan and Evermann 1898: 2260 (description). Robins 1958: 196 (habits in Florida Keys). Hildebrand, Chavez and Compton 1964: 123 (Alacran Reef, Yucatan). [B&R 1968]Böhlke

Barbulifer cauthoecus: Jordan, Evermann and Clark 1930: 446 (species name misspelled in check-list). Erdman 1956: 336 (Guanica, Puerto Rico; we did not examine the collection from Guajataca). [B&R 1968]

Barbulifer papillosus Eigenmann and Eiganmann 1888(1889): 70-72; holotype 26186 MCZ; type locality: Key West Florida; distinguished from ceuthoecus). [B&R 1968] [Eigenmann and Eigenmann 1888(1889)]

Barbulifer sp.: Caldwell and Caldwell 1964: 29 (Caledonia Bay, Panama).[ B&R 1968]

Barbulifer, species (in part): Caldwell 1966: 69 (Jamaica collection listed, some of them this species, others antennatus).[B&R 1968]


antennatus Böhlke and Robins 1968

Barbulifer antennatus Böhlke and Robins 1968: 126; Holotype ANSP 110642; Type locality: sta. 410, Bahamas, Grand Bahamas Bank, Exuma Cays, baylet on SE side of Warderick Wells Cay; to ca. 6 ft. [B&R 1968]

Barbulifer, species (in part): Caldwell 1966: 69 (some Jamaican collection listed are of this species, others are ceuthoecus). [B&R 1968]


mexicanus Hoese 1985

Barbulifer mexicanus Hoese 1985: 337-338; holotype SIO 61-248; type locality: Los Frailes, Baja California, Gulf of Mexico. [Hoese 1985]


pantherinus (Pellegrin 1901)

Gobiosoma pantherinum Pellegrin 1901: 165; syntypes MNHN 01-295; Type locality: Sargassum floating in the Gulf of California. Ginsburg 1933b: 17 (statement that pantherinum probably belongs to Barbulifer). [B&R 1968] [Hoese 1985- only source to list the type no.][Pellegrin 1901]

Gobiosoma spiriti-sancti Wade 1946: 219, pl. 30; holotype LACM 21556; type locality: Espiritu Santo Island, Gulf of California, Mexico). [B&R 1968] [Hoese 1985]


Genus Bollmannia


Bollmannia Jordan, 1890: 164(in: Jordan and Bollman 1890); type species Bollmannia chlamydes Jordan 1890: 164-165, by original designation.


chlamydes Jordan 1890:164-165; Lectotype USNM 93825, by subsequent designation of Ginsburg 1939: 58; Type locality: Panama Bay; Albatross station 2800, lat. 8051', long. 79031'30"; 7 fathoms. Note: Jordans original type, USNM 41158, does not match the description of the type and the jar contain two distinct species. [Ginsburg 1939]


boqueronensis Evermann and Marsh 1900: 356; holotype USNM 49366; type locality: sta. 6074 off Puerto Real, Puerto Rico, dredged by the Fish Hawk, in 8.5 fathoms of water. [Evermann and Marsh 1890]


communis Ginsburg 1942:364-365; holotype USNM 119873; type locality: Pelican station 77-1; 28059'N., 89029'W., 10 fathoms. [Ginsburg 1942]


eigenmanni Gamon

Gobius eigenmanni Gamon ??? USNM 231318


jeannae Fowler 1941: 95-96; holotype ANSP 69718; type locality: Key West, Florida.


litura Ginsburg 1935: 1-3; holotype USNM 93797; type locality: Caroline station 59. Samana Bay, Dominican Republic, 18-19 fathoms. [Ginsburg 1935]


longipinnis Ginsburg 1939: 60; holotype Bingham Oceanographic Collection 1690; Type locality: Angeles Bay, Gulf of California. [Ginsburg 1939]


macropoma Gilbert 1892:556; syntypes (6) SU 81 [Böhlke 1953]; type locality: Gulf of California just north of La Paz Bay, at Sta. 2996 (Albatross Exp.); 112 fathoms. [Gilbert 1892]


marginalis Ginsburg 1939: 59; holotype USNM 107284; type locality: Solango Island, Ecuador, 12 fathoms. [Ginsburg 1939]


ocellata Gilbert 1892:557; type not designated by Gilbert; Type locality: Northern part of the Gulf of California; Albatross Exp. Sta. 3031 and 3035, 30 and 33 fathoms. [Gilbert 1892] Lectotype USNM 107286 by subsequent designation of Ginsburg 1939: 59-60. Gilbert's original specimens were lost. His discription is based on a combination of three species. The only jar at the USNM labeled ocellata from the type locality was used to designate the lectotype. However, since it is not certain if Gilbert ever examined these specimens it may be a neotype and not a lectotype.


pawneea Ginsburg 1939: 60; holotype Bingham Oceanographic Collection 1689; Type locality: Perlas Islands, Panama Bay, lat.8029'40", long. 78052'30", 19-24 fathoms. [Ginsburg 1939]


stigmatura Gilbert 1892: 556; syntype SU 10 [Böhlke 1953]type USNM_____; type locality: northern part of the Gulf of California, Albatross Exp. Sta. 3016 and 3017, 76 and 58 fathoms [Glibert 1892]


umbrosa Ginsburg 1939: 59; holotype USNM 107290; type locality: Panama Bay, Albatross station 2804, lat. 8016'30", long. 79037'45", 47 fathoms. [Ginsburg 1939]



Genus Chriolepis


Chriolepis Gilbert, 1892:557;Type species Chriolepis minutillus Gilbert 1892:558 by original designation. [Gilbert 1892]


minutillus Gilbert 1892: 558; holotype USNM 48261 [Gilmore 1979] type locality: Albatross Exp. Sta. 2825, Gulf of California [Findley 1975] [Gilbert 1892]

benthonis Ginsburg 1953:22 (type locality: off Progreso, Yucatan, Mexico) holotype (unique) USNM 47641


cuneata Bussing 1990:113-115. (type locality: Isla San Jose, Islas Murcielagos, Costa Rica) depth 10-14 meters. holotype LACM 4428-1; paratypes also at LACM


dialepta Bussing 1990:115-117. (type locality: East side Isla del Coco, 300 m SW Cabo Desubierta, in 20 meters depth); holotype LACM 44821-4; paratypes also at LACM


Chriolepis walkeri, Findley 1983, unpublished Ph.D. dissertation, Univ. of Arizona, 193p.



fisheri Herre 1942: 297; holotype _____ not listed in publication; type locality: Barbados, depth 45 fathoms. [Herre 1942]

lepidotus Findley 1975:94-97; holotype USNM 211456; type locality: Malpelo Island, Colombia, east side of island, depth ca. 10 meters, above the coral zone, cobble bottom with some calcareous sand and boulders, very sparse algae growth. [Findley 1975]


tagus Ginsburg 1953:21-22; holotype USNM 123232; type locality: Tagus Cove, Albemarle Island, Galapagos Archipelago, 10-18 fathoms. [Ginsburg 1953]



vespa (Hastings & Bortone) 1981: 428-432; holotype USNM 221523; type locality: northeastern Gulf of Mexico, 28024'N, 85015'W, R/V Bellows (cruise Bureau of Land Management, BLM 15, station III-C, 183 m, 26 July 1975. [H&B 1981]


Eleotriculus (subgenus) Ginsburg 1938: 111; type species Chriolepis zebra Ginsburg 1938 by original designation. [Ginsburg 1938]


zebra Ginsburg 1938: 109-111; holotype in Hancock Coll.; Type locality: San Gabriel Bay, Espiritu Santo Island, Gulf of California, shore collection. [Ginsburg 1938]


Genus Eleotrica


Eleotrica Ginsburg 1933a: 10; Type species: Eleotrica cableae Ginsburg 1933, by original designation.

  Alepidoeleotris Herre 1935:413; synonym of Eleotrica Birdsong and Robins 1995:682


cableae Ginsburg 1933a:11-12; type USNM 65517; Type locality: Chatham Island, Galapagos Archipelago [G. 1933]

Alepidoeleotris tigris Herre (1935:413) synonomized by Myers (1936:129) without checking the specimen; Eschmeyer and Bailey (1990:19) without discussion; Birdsong and Robins (1995:682) with discussion.


Genus Enypnias


Enypnias Jordan and Evermann 1898: 2233-2234. Type species: Gobius seminudus Gunther 1861, by original designation; proposed as a subgenus of Garmannia. [B&R 1968]


seminudus Günther) 1861a

Gobius seminudus Günther 1861:a 372; Type ____ (number and location of specimen not listed in original publication); Type locality: west coast of Central America. 1861b: 554 (description); 1862:328 (repeat of original description). [B&R 1968] [Günther 1861a]


aceras Ginsburg 1939a: 61; Holotype USNM 107298; Type locality: Miraflores Locks, lower chamber, Pacific slope, Panama Canal. [Ginsburg 1939a][B&R 1968]

Enypinas aceras: Hildebrand 1939: 43 (genus misspelled; 127 specimens recorded from upper and lower chambers of Miraflores Locks). [B&R 1968]


Genus Evermannichthys


Evermannichthys Metzelaar, 1919: 139-140 figs 39,40. Type species: Evermannichthys metzelaari Hubbs 1923:2 (by monotypy).

Radcliffella Hubbs, 1921: 2; Type species Garmannia spongicola Radcliffe 1917, by original designation. [B&R 1969]


spongicola (Radcliffe 1917)

Garmannia spongicola Radcliffe, 1917: 423-425, fig. (unnumbered); Holotype USNM 77501; Type locality: North Carolina, 1 mile SE of fishing buoy, 34019'N, 76048' W in 14: fms., from sponge. Hubbs, 1921: 2 (made type of new genus Radcliffella). Kousmans, 1931: 72-73 (characters, nomenclature; retained in Evermannichthys). [B&R 1969][Radcliffe 1917]

Evermannichthys spongicola: Hubbs, 1923: 1-2 ( spongicola placed in Evermannichthys Metzelaar and compared to E. metzelaari). Hildebrand, 1941: 229 (North Carolina; compiled). Delsman, 1941: 76 (9 specimens from near Tampa). Springer, 1961: 482 (records near Tampa Bay, compiled). Moe et al., 1966: 75 (4 localities on West Coast of Florida south of Tampa Bay). [B&R 1969]

Evermannichthys metzelaari (misidentifications): Pearse, 1932: 119 (Tortugas; reported from sponge, Speciospongia vespara [sic]). Longley and Hildebrand, 1941: 229-230 (in part; descriptive data on 3 specimens from Tortugas). Pearse, 1950: 151 (taken at Tortugas; compiled). Briggs, 1958: 288 (in part; Florida distribution). Duarte-Bello, 1959: 122 (reference to eastern Florida only). Bullis and Thompson, 1965: 56 (Gulf of Mexico records). [B&R 1969]


bicolor Thacker 2001

Evermannichthys bicolor Thacker 2001:2-4, fig 1; Holotype LACM 54100-3; Type locality: Navassa Island, Caribbean Sea, rotenone station in a flat sand habitat with scattered rocks, gorgonians, and sponges at a depth of 27-30 feet.

convictor Böhlke and Robins 1969

Evermannichthys convictor Böhlke and Robins 1969: 11-13, fig. 2;by original designation; Holotype ANSP 111862; Type locality: sta. TS-7, Bahamas, Graet Bahama Bank, Green Cay (E of southern Andors ISland), W side of cay, 100 yards S of sand spit at far NW end; 45 ft.; 10 Jan. 1968. [B&R 1969]


metzelaari Hubbs 1923

Evermannichthys spongicola Metzelaar, 1919: 139-140, figs 39-40 (description; type locality; Curacao); 1922: 141 (generic diagnosis). [B&R 1969]

Evermannichthys metzelaari Hubs, 1923:1-2 (substitute name for E. spongicola, preoccupied as result of referral to Evermannichthys of Garmannia spongicola Radcliffe). Beebe and Tee-Van, 1928: 223-224 (characters, ecology, Haitian records). Koumans, 1931: 72-73 (nomenclature, discussed under the names Evermannichthys spongicola and E. metzelaari plus two, never used, manuscript manes Evermannella spongicola and Diogenides cavitarius). Longley and Hildebrand, 1941: 229-230 (in part; references to Dutch West Indies only). Briggs, 1958: 288 (in part; Lessrer Antilles only). Duarte-Bello, 1959: 122 (in part, Lesser Antilles reference only). Böhlke and Chaplin 1968:627 (characters, Bahaman record, distribution, figure). [B&R 1969]


silus Böhlke and Robins 1969

Evermannichthys silus Böhlke and Robins 1969: 10-11,fig. 1; by original designation; Holotype ANSP 111865; Type locality: sta. TS-71, Bahamas, Samana (Atwood) Cay, E of the Graet Bahama Bank, middle of W half of S side of cay; 90 ft. 27 Jan. 1968. [B&R 1969]


Genus Ginsburgellus


Ginsburgellus Böhlke and Robins 1968: 140-141. Type species Gobiosoma novemlineatum Fowler 1950 = Ginsburgellus novemlineatus, the only species in the genus. [B&R 1968]


novemlineatus (Fowler 1950)

Gobiosoma novemlineatum Fowler 1950: 89, figs. 44-46. holotype ANSP 71778; Type locality: Caribbean Sea, San Andres [St. Andrews Island] off coast of Nicaragua.  Erdman 1956: 335 (Puerto Rico; color; association with sea urchin, Echinometra). [B&R 1968][Bohlke, E.B. 1984, for holotype number]


Genus Gobiosoma


Gobiosoma Girard 1858: 169 (type species: Gobiosoma molestum Girard 1858 = Gobiosoma bosc Lacepède 1800, by subsequent designation by Bleeker, 1874: 310)


Subgenus Gobiosoma


Gobiosoma Girard 1858: 169 (type species: Gobiosoma molestum Girard 1858 = Gobiosoma bosc Lacepède 1800, by subsequent designation by Bleeker, 1874:310)


                        Dilepidion Ginsburg 1933a: 17; Type species: Gobiosoma ginsburgi Hildebrand and Schroeder 1928, by original designation.[Ginsburg 1933a]


bosc (Lacepède, 1798)

Gobius bosc Lacepède 1800:555 (Charlston, S.C., from a manuscript by Bosc.; as restricted by Hildebrand and Schroeder.)[Gins. 1933b]

Gobius alepidotus Bloch and Schneider 1801 (Apparently based on Lacepède's account)[Gins. 1933b]

Gobius viridipallidus Mitchil 1815 (New York; as restricted by Cuvier and Valenciennes)[Gins. 1933b]

Gobiosoma molestum Girard 1858 (Texas; type reexamined; based on a specimen of the present species). Girard; 1859(Texas, based on same specimen as preceding record). Günther; 1861 (Texas, after Girard). Jordan and Gilbert, 1882 (Texas after Girard).[Gins. 1933b]

Gobiosoma alepidota Uhler and Lugger; 1876 (Worcester County, Maryland). [Gins. 1933b]

Gobiosoma bosci Bean 1889 (Somers Point and Ocean City, N.J.). Bean 1890(Long Island, New York). Bean, 1892 (Cape Charles, Va.). Evermann and Kendall, 1894 (Galveston and Corpus Christi, Texas). Smith and Bean (in part), 1898 (Potomic River at Gunston Wharf, VA). Evermann, 1898, 1899 {could be 1900 ref. see bib.}, (Baldwin Lodge, Miss.). Bean, 1898 (Long Island, many localities, some of the specimens reexamined). Jordan and Dickerson, 1908 (Tampico, Mexico). Hildebrand and Schroeder, 1928 (Chesapeak Bay, many localities).[Gins. 1933b]

            Distribution: From Long Island, NY to Tampico, Mexico[Gin.33]; it is absent from southeastern Florida and the Keys [B&R 68]

Biology: Dahlberg, M.D. et al 1973 (ecology and reproduction - Georgia; compared with ginsburgi)


ginsburgi Hildebrand and Schroeder 1928:324 type USNM 87655 Type locality Chesapeake Bay. [H&S, 1928]

Biology: Dahlberg et al 1973( reproduction and ecology in Georgia; compared with bosci)


heterolepidotum Hoese 1971:76 (unpublished MS name)Type SIO 67-274 (formerly UA 65-33) Type locality Punta Peñasco, Sonora, Mexico. Conglomerate reef, midtidal pool. Collected July 27, 1965.


longipala Ginsburg 1933a:18; holotype USNM 86158; Type locality: Boca Grande, Florida, 2 April 1917; taken by steamer Grampus. [Ginsburg 1933a]


Subgenus Austrogobius


Austrogobius de Buen 1950: 122 (name introduced); 1951: 64-68 Type species: Gobiosoma parri Ginsburg 1933, by monotypy. [B&R 1968]


parri Ginsburg 1933b: 44-46; Holotype: USNM 93177; Type locality: Pocitos, Uruguay. Fowler 1943: 330 (five specimens from Uruguay; eight dorsal spines in error). Dawson 1963: 585 (in subgenus Dilepidion, name only). [B&R 1968]

Gobius sp.: Barattini 1942: 13 (coast of Uruguay). [B&R 1968]

Austrogobius parri: de Buen 1950: 122 (synonymy; announcement of genus); 1951; 63-68, pl. 1 (made type of new genus Austrogobius; redescription, illustrated). [B&R 1968]

Garmannia parri: Robins 1960: 284 (Austrogobius synonymized with Garmannia). Robins and Bohlke 1964: 5 (retained in Garmannia). [B&R 1968]


grosvenori (Robins) 1964:399-402; holotype ANSP 101166; Paratypes UMML 14380; Type locality: Boca Chita Pass, Dade Co., FL; Cervigon 1966: 743-745, fig. 313 (three specimens from Cubagua Island, Venezuela). Caldwell 1966: 70 (specimens from Palisadoes, Jamaica). [B&R 1968][Robins 1964]


hemigymnum (Eigenmann and Eigenman) 1888(1889)

Gobius hemigymnus Eigenmann and Eigenmann 1888(1889): 66-67; holotype MCZ no. 27124; Type locality: West Indies, exact location unknown). [B&R 1968] [Eigenmann and Eigenmann 1888(1889)]

Garmannia hemigymna: Ginsburg 1933b: 57-58 (placed in new subgenus Risor). Fowler 1940: 791 (two specimens from Rio de Janeiro). Fowler 1941: 177 (Rio de Janeiro; in check list). Fowler 1951: 30-31 (redescription on basis of two specimens from Rio de Janeiro). Robins 1960: 284 ( removed from genus Risor; characters summarized). [B&R 1968]

Garmannia mediocricula Ginsburg 1942: 365-366 Type locality: Rio de Janeiro; description, based on same material as in Fowler's references above). Robins 1964: 402 (compared with grosvenori). [B&R 1968?] [Ginsburg 1942]


spilotum (Ginsburg) 1939a:62

Garmannia spilota Ginsburg 1939a: 62; holotype USNM 81828; Type locality: Colon, Panama, in tidepools. Robins 1960: 284 (compared to Garmannia saucra). Robins 1964: 402 (compared to grosvenori).[Ginsburg 1939a][B&R 1968]


Subgenus Garmannia


Garmannia Jordan and Evermann, in Jordan et al. 1895: 497 Type species: Gobius paradoxus Gánther 1861:372, by original designation.[B&R 1968]

Gerhardinus Meek and Hildebrand 1928: 889 Type species Gerhardinus nudus Meek and Hildebrand 1928, by original designation.[B&R 1968] [Meek and Hildebrand 1928]

Gobiolepis Ginsburg 1944: 379 Type species: Garmannia hildebrandi Ginsburg 1939a, by original designation.[B&R 1968][Ginsburg 1944]

Gobiohelpis Ginsburg 1944: 380 Type species: Garmannia spes Ginsburg 1939a:69, by original designation.[B&R 1968][Ginsburg 1939a][Ginsburg 1944]

Gobiculina Ginsburg 1944: 380 Type species: Garmannia homochroma Ginsburg 1939:62, by original designation. [B&R 1968][Ginsburg]


chiquita (Jenkins and Evermann) 1889

Gobius chiquita Jenkins and Evermann 1889: 146; Holotype USNM 39634; Type locality: Guaymas, Mexico; spines erroneously listed as six. [B&R 1968]


hildebrandi (Ginsburg 1939a)

Garmannia hildebrandi Ginsburg 1939a: 61-62; holotype USNM 107297; Type locality: Gatun Locks, upper lock, Atlantic side of Panama Canal; also from Pedro Miguel Locks, Pacific side. Hildebrand 1939: 30, 33 (Gatun and Pedro Miguel Locks, report on the type material). Fowler 1944: 518 (compiled). Ginsburg 1942: 366 (compared with Garmannia mediocricula). Ginsburg 1944: 379 (made type of subgenus Gobiolepis; characters). Ginsburg 1945: 136-137 (caudal-ray counts). Robins 1960: 284 (compared with Garmannia saucra). Robins 1964: 402 (compared with Garmannia grosvenori).[Ginsburg 1939a] [B&R 1968]


homochroma (Ginsburg) 1939a

Garmannia homochroma Ginsburg 1939a: 62-63; Holotype USNM 107296; Type locality: Pedro Miguel Locks Pacific slope of Panama. Hildebrand 1939: 33 (listed ten specimens as type material from Pedro Miguel Locks). [B&R 1968][Ginsburg 1939a]


nudum (Meek and Hildebrand) 1928

Gerhardinus nudus Meek and Hildebrand 1928:889, pl. 88; holotype USNM 81834; Type locality: tidepools, Panama City, Pacific Coast of Panama; type of genus Gerhardinus. Brittan 1966: 132 (Mata Limon estuary, Costa Rica). [B&R 1968] [Meek and Hildebrand 1928]

Gobiosoma nudum: Ginsburg 1933b: 46-47 (Gerhardinus placed as subgenus of gobiosoma; characters, relationships, range extended to Peru). Hildebrand 1939: 43 (lower chambers, Miraflores Locks, Panama). Fowler 1944: 518 (compiled). Rubinoff 1963: 140-145 (distribution in Panama). [B&R 1968]


paradoxum (Günther) 1861a

Gobius paradoxus Günther 1861a: 372; Type _____ (not listed in original publication); Type locality: western coast of Central America; dorsal spines erroneously listed as six; 1861b: 549-550 (description); 1862: 327 (repeat of original description). Gilbert and Starkes 1904: pl. 38 (good illustration of type species). [B&R 1968] [Gánther 1861a]


robustum Ginsburg 1933a:15-16; Type USNM 92802; Type locality Laguna Madre, near Corpus Pass, Texas [Gin. 1933]

Biology: Springer & McErlean 1961: 73-75; Springer & McErlean 1961; Dawson 1966: 384-392; Zavala Garcia 1988[1989] (early life history & larval distribution, Mexico)

Distribution: Gulf Coast from southern Texas to Key West, FL and alopng the Atlantic coast to Miami, northward to Cape Canaveral, FL [B&R 68]

ERRORS: Record from Bahia, Brazil most likely G. parri[B&R 68]; record from Jamaica by Caldwell 1966:70 is Psilotris alepis Ginsburg [B&R 68]


schultzi (Ginsburg) 1944

Garmannia schultzi Ginsburg 1944: 375-378; holotype USNM 121546; paratypes (6) USNM 121547; Type locality: Venezula, Lake Maracaibo, 7 km S of Maracaibo City. Schultz 1949: 185 (collections from Maracaibo region of Venezula). Robins 1960: 280 (contrasted with Garmannia saucra).[B&R 1968][Ginsburg 1944]


spes (Ginsburg) 1939a

Garmannia spes 1939a: 62; holotype USNM 107299; Type locality: Canal Zone, dry dock, Mt. Hope. Ginsburg 1942: 366 (contrasted with Garmannia mediocricula). Ginsburg 1944: 377-378, 380 (data on specimens from Venezula; made type species of new subgenus Gobiohelpis). Fowler 1044: 518 (compiled). Schultz 1949: 185 (collection from near Sinamaica, Venezula). Caldwell et al. 1959: 28 (recorded from Costa Rica). Robins 1960: 284 (contrasted with Garmannia saucra). [B&R 1968][Ginsburg 1939a]


Garmannia hildebrandi: Caldwell 1966: 70 (misidentification, collection from Palisadoes, Jamaica). [B&R 1968]


yucatanum Dawson 1971: 433-437; holotype USNM 205368; type locality: Mexico, Quintana Roo, south side of harbor of Cd. Chetumal; 1 meter.


Subgenus Gobiolepis

This subgenus is included because of the new species, which are being described and placed in it.  Several other species above also belong here but are being retained in their present subgenera until the manuscripts appear in press.


Gobiolepis Ginsburg 1944

Originally defined as a subgenus of Garmannia by Ginsburg, Type species Garmannia hildebrandi Ginsburg 1939. Now defined by Hoese '71 (unpublished thesis) as a subgenus of Gobiosoma and to include the following species: G. hildebrandi, G. birdsongi (sp. nov.), G. chiquite, G. zacae (sp. nov.), G. grosvenori, G. homochroma.


birdsongi  Hoese 1971(unpublished MS name):Holotype ANSP (formerly UMML 23466) type locality a mud-covered, rocky flat and stream at base of spillway in Miraflores Locks of the Panama Canal.


zacae  Hoese 1971(unpublished MS name): Holotype SU 60374; Type locality - Panama, Bahía Honda.


Subgenus Elacatinus


Elacatinus Jordan 1904: 542; Type species: Gobiosoma oceanops Jordan 1904, by original designation and by monotypy. This subgenus frequently has been misspelled in the literature, Elecatinus and Lactinius being the most common misspellings  [B&R 1968]


oceanops Jordan 1904

Elacatinus oceanops Jordan 1904: 542, pl.2, fig.3 Holotype Stanford Nat. Hist. Mus. 8365, cotype 2757; Type locality: Garden Key, Tortugas, Florida; habitat on coral heads at depths of 3-8 ft.(original description) [Feddern 1967]

Gobiosoma oceanops: Ginsburg 1933: 2, 19, 21-22. Briggs 1958: 288 (range). Duarte-Bello 1959: 123 (range).[Feddern 1967] Hildebrand, Chavez and Compton 1964: 124 (Alacran Reef, off Yucatan). [B&R 1968]

Lactinus oceanhops: Fischer 1955: 36-37, 70 (name misspelled; general and spawning behavior in aquaria) [Feddern 1967] Straughan 1956: 157 fig. (popular notes on species in the aquarium) [B&R 1968]

Lactinius oceanops: Straughan 1959: 140-144, fig. 62 & 63 (name misspelled). Straughan 1964: 165-169, fig. 21, 56, 72, 73, & 75 (spawning in aquarium) [Feddern 1967]

Elecatinus oceanops: Herold 1961: 235, pl. 100 (name misspelled). Limbaugh 1961: 43 (parasite picking behavior). [Feddern 1967]

?Elacatinus oceanops: Wikler 1962: 201-205 (general and spawning behavior; description and fig. of egg and larve). [Fedder 1967]

Elecatinus oceanhops: Menten 1962: 246, figs. (misspelled at beginning of paper, notes on spawning in aquarium). [B&R 1968]

Elacatinus oceanops: Schroeder 1965: 110-112 (Alligator Reef, Florida Keys; parasite removing habits). Eibl-Eibesfeldt 1967: fig. 1. Starck and Davis 1966: 340, figs 21 & 22 (inactive at night). [B&R 1968]

not Elacatinus oceanops: Breder 1927: 83 (West Indies). Fowler 1944: 451 (Bahamas). Eibl-Eibesfeldt 1955: 207-209, figs. 5-7 (description of parasite removing at Bonaire, Dutch West Indies) [Feddern 1967]

not Gobiosoma oceanops Erdman 1956: 335 (Jamaica; Haiti; Puerto Rico). [Feddern 1967]


chancei Beebe & Hollister 1933

Gobiosoma chancei Beebe and Hollister 1933: 87-88, fig. 17; holotype USNM 170955; type locality: Grenada, British West Indies, St. George's Bay. [Beebe &Hollister 1933] Ginsburg 1933b: 25 (synonymized in error with G. horsti). [B&R 1968]

Gobiosoma horsti: Herre 1942: 14[298] (Hastings, Barbados, taken from a sponge). Cervigon 1966: 740-741 (in part: figure and account based on chancei, reference on horsti, and Tortugas part of range based on xanthipora). [B&R 1968]


atronasum Böhlke & Robbins 1968

Gobiosoma (Elacatinus) astronasum Bohlke and Robins 1968: 88-90, fig. 6; holotype UF 15427; Type locality: Bahama Islands, at the eastern edge of the Great Bahama Bank. [B&R 1968]


evelynae Böhlke & Robbins 1968

Gobiosoma  (Elacatinus) evelynae Böhlke and Robins 1968: 103-109, fig. 10; holotype ANSP 110559; Type locality: sta. 496, Bahamas, Great Bahama Bank, Nassau vicinity, Green Cay (N of Rose Island), ca. 1/4 mi. NNW of Cay; ca. 50 ft. [B&R 1968]

Elacatinus oceanops: Eibl-Eibesfeldt 1957: 38 (Bonaire; discussion of cleaning, with drawing of six gobies cleaning an Epinephelus striatus, including inside of mouth and under operculum; probably, although not certainly, evelynae). [B&R 1968]


figaro Sazima, Moura and Rosa 1996

Elacatinus figaro Sazima, Moura and Rosa, 1996:33-38, figs. 1-6; Holotype MZUSP 50859; Type locality: Pedra do Navio, Iiha Anchieta, Ubatuba, Sao Paulo, Brazil, depth 5 m, 02 May 1996.


genie Böhlke & Robbins 1968

Gobiosoma (Elacatinus) genie Böhlke & Robins 1968: 109-114, fig. 11; holotype ANSP 110574; Type locality: sta. 243, Bahamas, Great Bahama Bank, Nassau vicinity, between North Cay and the small cay just off Long Cay; 6-13 ft. [B&R 1968]

Elacatinus oceanops: Breder 1927: 83 (Saddle Rock and Washerwoman Cut, Bahamas; probably of this species). Fowler 1944: 451 (in checklist; based on Breder 1927 ref.). Feder 1966: 342-343, 360 (cleaning habits in the Bahamas; identification uncertain but probably this species). [B&R 1968]


horsti Metzelaar 1922

Gobiosoma horsti Metzelaar 1922: 139, fig. 2; Lectotype Zool. Mus. Amsterdam 446; Type locality: Caracasbay, SW part of Curacao, from sponge. Beebe and Tee-Van 1928: 224, fig. 17 (specimens from Port-au-Prince Bay, Haiti; descriptive information and habitat based on more than one species). Jordan, Evermann and Clark 1930: 446 (compiled). Beebe and Hollister 1931: 88 (compared to chancei). Ginsburg 1933b: 22-25 (in part: chancei incorrectly included in synonymy). Fowler 1944: 472 (in part: Grenada record refers to chancei, Tortugas record to xanthiprora). Fowler 1952: 109 (Haiti; compiled). Briggs 1958: 288 (compiled; in part: Tortugas record based on xanthipora). [B&R 1968]


illecebrosum Böhlke & Robbins 1968

Gobiosoma (Elacatinus) illecebrosum Böhlke & Robins 1968: 114-116, fig. 12; holotype ANSP 110693; Type locality: sta. WAS-CARIB-8, Yucatan, Banco Chinchorro, around Cay Norte; 15-20 ft. [B&R 1968]


louisae Böhlke & Robbins 1968

Gobiosoma (Elecatinus) lousiae Böhlke and Robins 1968: 90-94, fig.7; holotype ANSP 110691; Type locality: sta. 14 (G64-43), W side of Grand Cayman Island at dropoff, offshore from N side of Georgetown; 100-115 ft. [B&R 1968]


prochilos Böhlke & Robbins 1968

Gobiosoma (Elacatinus) prochilos Böhlke and Robins 1968: 97-100, fig.9; holotype ANSP 110628; Type locality: sta. TE-57, St. Barthelemy, Port de Gustavia, off the rocky cliffs just S of Anse Galet; 40-45 ft. [B&R 1968]


randalli Böhlke & Robbins 1968

Gobiosoma (Elacatinus) randalli Böhlke and Robins 1968: 94-97, fig. 8; holotype ANSP 110672; Type locality: sta TE-25, Grenadines, St. Vincent Island, N end of Anse Mahaut, just S of Cumberland Bay; 20-25 ft. [B&R 1968]


tenox Böhlke & Robbins 1968

Gobiosoma (Elacatinus) tenox Bohlke and Robins 1968: 83-85, fig. 4; holotype ANSP 110688; Type locality: sta. TE-42, Dominica, Prince Rupert Bay, Portsmouth; 50ft. [B&R 1968]


xanthiprora Bohlke & Robbins 1968

Gobiosoma (Elacatinus) xanthipora Böhlke and Robins 1968: 85-87, fig.5; holotype ANSP 110898; Type locality: Florida, Monroe Co., 1/4 mi. SE of Alligator Reef Light; about 75 ft. Paratypes: USNM 118105: Tortugas; UMML 8312: Jamaica. [B&R 1968]

Elacatinus horsti: Longley and Hildebrand 1941: 227 (in part: synonymy given therein does not apply to xanthipora, but Tortugas specimens described are that species). [B&R 1968]

Gobiosoma horsti: Briggs 1958: 288 (in part: Tortugas record). Bailey et al. 1960: 36 (name only; based on Tortugas record). Caldwell 1966: 70 (specimen from Lime Cay, Jamaica. [B&R 1968]


Subgenus Tigrigobius


Tigrigobius Fowler 1931: 401 Type species Gobiosoma macrodon Beebe and Tee-Van 1928, by original designation. [B&R 1968]

Gobicula Ginsburg 1944: 379. Type species: Garmannia gemmata Ginsburg 1939, by original designation.[Bohlke & Robbins 1968]


macrodon Beebe & Tee-Van 1928:226; holotype originally no. 7462 in the Tropical Research Collection of the New York Zoological Society now at USNM 170896; type locality: Port-au-Prince Bay, Haiti. [Robins 1958] [Beebe and Tee-Van 1928]see: Robins 1958:195-196 for syn. & distribution

Springer & Woodburn 1960: 72 range

digueti (Pellegrin, 1901) Listed as Elacatinus digueti in Thomson et al 1979 where it is synonomized with brocki [Thomson 1979, p203]

Gobiosoma brocki Ginsburg 1938: 58-59; holotype Stanford University No. 32970; Type locality: Los Frailes near Cape San Lucas, Baja CA. Fowler 1944: 518 (compiled). [B&R 1968][Ginsburg 1938]


dilepis (Robins & Bohlke 1964)

Garmannia (Tigrigogius) dilepis Robins and Bohlke 1964: 3-6, fig 2; holotype ANSP 100815; Type locality: Sta. 589, coral head off large bay at NW end of Conception Island, Bahamas, 25 ft.. [B&R 1968][R&B 1964]


gemmatum (Ginsburg 1939a)

Garmannia gemmata Ginsburg 1939a:3-5, fig. 2; holotype USNM 107291; Type locality: Old Providence Island, Caribbean Sea. Herre 1942: 14 (recorded from Barbados). Ginsburg 1944: 379 (placed in new subgenus Gobicula). Bohlke and Robins 1960: 92-93 (Gobicula synonymized with subgenus Tigrigobius of Garmannia; dentition and squamation discussed). Robins 1960: 284 (contrasted with Garmannia saucra). [B&R 1968][Ginsburg 1939]

Garmania gemmata: Schmitt and Schultz 1940:8 (generic name mispelled; type specimens listed only).[B&R 1968]



inornatus Bussing 1990:109-110 (type locality: Isla Cocinero, Islas Murcielagos, Costa Rica (10051'N, 85054'W) depth 9 meters. holotype LACM 32493-43; paratypes also at LACM.

Gobiosoma brocki (not Ginsburg) Böhlke and Robins (in part) 1968:67-69 (Isla Gorgona, Columbia; not including Panama specimens)

Gobiosoma sp., Burgess & Axelrod, 1984: 2109 (color plate of freshly preserved specimen)

Elacatinus sp. Lopez & Bussing, 1982.


janssi Bussing 1981: 251-256, fig. 1. (type locality: Bahia Herradura just south of the Gulf of Nicoya (9038'45"N, 84040'55"W), Pacific coast of Costa Rica) Holotype LACM 32524-45; paratypes UCR 618-23, UCR 628-11, LACM 32509-37, UCR 638-3 (other paratypes listed in manuscript)



limbaughi Hoese and Reader 2001

Elacatinus limbaughi Hoese & Reader 2001 More info to follow soon


multifasciatum Steindachner 1876

Gobius lineatus Poey 1861 (a homonym of Gobius lineatus Jenyns, 1842)[Gins. 1933b]

Gobiosoma multifasciatum Steindachner 1876:Sitzbar. k. k. Ak. Wiss. Wien 74 (abt.1): 231 and 238 (Ichthyol. Beit. 5: 183 and 190); Type locality: St. Thomas; Barbados and Bartholomew, Lesser Antilles. [Gins.1933b]

see: Ginsburg 1933b for additional references.


nesiotus Bussing 1990:110-112; (type locality: East side of Isla del Coco, 300m SW of Cabo Descubierto; depth 20 meters);holotype LACM 44821-3; paratypes also at LACM.


pallens (Ginsburg) 1939a

Garmannia pallens Ginsburg 1939a:63; holotype USNM 107327; Type locality: Barahona Harbor, Santo Domingo, 2-4 ft. [Ginsburg 1939a][B&R 1968]

see: Bohlke & Robins 1960 for summary & literature


puncticulatum Ginsburg 1938

Gobiosoma punticulatum Ginsburg 1938: 57-58; holotype Stanford University No. 32963; Type locality: Los Frailes, near Cape San Lucas, Baja California. Fowler 1944: 518 (in check-list only). [B&R 1968][Ginsburg 1938]

Gobiosoma rubrifrons Fowler 1944: 282, 383, fig. 237 (type locality: Saboga, Perlas Islands, Gulf of Panama). [B&R 1968]


saucrum (Robins 1960)

Garmannia saucra Robins 1960: 281-285, 1 fig.; holotype ANSP 92985; Type locality: South Cay, Jamaica, 3 miles south of Port Royal, north side of cay in 3-5 ft. of water. Caldwell 1966: 70 (compiled) [B&R 1968] [Robins 1960]


zebrella (Robins) 1958

Garmannia zebrella Robins 1958:192-194; holotype ANSP 53387; type locality: Monos Island, Trinidad. [Robins 1958]


Genus Gobulus


Gobulus Ginsburg, 1933a: 12. Type species: Gobulus crescentalis (Gilbert) = Gobiosoma cresentalis Gilbert, 1892 {see note in bibliography on Gilbert}


crescentalis (Gilbert) 1892:557

Gobiosoma cresentalis Gilbert, 1892:557 [Gin. 1933a] Type locality: Albatross Exp. Sta. number 2825; Gulf of California, lat. 24022'15"N., long. 110019'15"W., 7 fathoms [Gilbert 1892]

Gobiosoma crescentale Jordan and Everman 1898:2259 U.S. Nat. Mus. Bull. 47, pt. 3 (erroneously said to have been taken in 79 fathoms) [Gin. 1933a]

Gobiosoma cresentale Pellegrin 1901:162 Bull. Mus. Hist. Nat. Paris, vol 7, (Gulf of California) [Gin. 1933a]

Gobiosoma crescentale Osburn and Nichols 1916:175 Bull. Amer. Mus. Nat. Hist., vol. 35 (Agua Verde Bay, Gulf of California)[Gin. 1933] [Osburn and Nichols 1916]


birdsongi Hoese and Reader 2001 more info soon


hancocki Ginsburg 1938: 118-119; holotype in USNM; Type locality: Secus Island, Panama; shore, coral tide flat. [Ginsburg 1938]


myersi Ginsburg 1939a:63; holotype USNM 107283; type locality: Gulf of Mexico, off Cape Sable; Albatross station 2374, lat. 29011'30", long. 85029', 26 fathoms. [Ginsburg 1939a] Note: according to Hastings 1983 the type locality is the extreme northeastern Gulf of Mexico. Ginsburg had the correct latitude and longitude but listed the wrong location for those marks. [Hastings 1983]


Genus Gymneleotris


Gymneleotris Bleeker 1874: 304 Type species: Eleotris seminudus Gánther 1864, by original designation. [B&R 1968]


seminudus (Gánther 1864)

Eleotris seminudus Gánther 1864: 24, pl. 4, fig. 2, 2a; Type _____ (type not listed in publication); Type locality: Pacific coast of Panama. [B&R 1968] [Gánther 1864]

Eleotris seminuda: Gánther 1869: 441 (redescription of type specimen). [B&R 1968]

Gymneleotris seminuda: Jordan and Eigenmann 1887: 484 (listed in review of North American Gobies). [B&R 1968]

Gymneleotris seminudus: Jordan and Evermann 1898: 2204 (description after Gánther). [B&R 1968]


Genus Microgobius


Microgobius Poey, 1876; Type species: M. signatus Poey, 1876: 168-169, pl. 8, fig.3 (type locality Cuba; syntypes MCZ 13127) [Birdsong 1981, all references unless otherwise stated]


Xenogobius weberi Metzelaar, 1919: 140-141 (type locality Curacao reef waters). Longley, 1932: 300 (junior synonym of Microgobius signatus Poey).[Birdsong 1981]

Zalypnus Jordan and Evermann, Check List, 1895(96?), 459 (Gobius emblematicus Jordan and Gilbert)[Jordan 1930]


signatus Poey, 1876; 168-169, pl. 8; fig. 3; syntypes MCZ 13127; type locality: Cuba

Xenogobius weberi Metzelaar, 1919: 140-141 (type locality Curacao reef waters). Longley, 1932: 300(junior synonym of Microgobius signatus Poey)


brevispinis Ginsburg 1939: 56 (type locality Balboa, Panama; holotype USNM 81842)

Gobius emblematicus Jordan and Gilbert 1882a: 330-331 (in part; see Birdsong 1981 for details)

Microgobius emblematicus Jordan and Eigenmann 1886: 503-504 (in part)


carri Fowler, 1945: 322 (type locality Sanibel Island, Fla; holotype ANSP 71187)[Birdsong 1981]


crocatus Birdsong 1968: figs. 1,2 (type locality Pacific coast of Panama near Miraflores Locks; holotype USNM 202587)


Microgobius miraflorensis Chirichigno Fonseca, 1968: 454, fig. 25 (Puerto Pizarro, Peru; based on male specimens; illustrated in error)

Microgobius tabogensis Chirichigno Fonseca, 1968: 457, fig 26 (Puerto Pizarro, Peru; based on female specimens; illustrated in error)


curtus Ginsburg 1939: 56 (type locatity Salada, Guayaquil, Ecuador, in mangrove swamp; holotype USNM 107292)

Microgobius emblematicus: Chirichigno Fonseca, 1968: 455, fig. 24 (Puerto Pizarro, Peru; illustration in error)


cyclolepis Gilbert 1891: 74 (type locality: northern Gulf of California, Albatross sta. 3020)

Zalypnus cyclolepis: Jordan and Evermann, 1896: 459 (assigned to Zalypnus)


emblematicus (Jordan and Gilbert) 1882: 330-331 (type locality Bay of Panama;[type series mixed according to Birdsong 1981] Neotype designated by Ginsburg 1939a:56. Neotype Stanford University Coll. 33208


M. cyclolepis Gilbert 1891 [Jordan, 1930][not according to Birdsong 1981]

Gobius emblematicus Jordan and Gilbert, 1882a

Lepidogobius emblematicus Jordan and Gilbert 1882c

Microgobius emblematicus Jordan and Evermann 1886: 503-505 (new combination)

Zalypnus emblematicus Jordan and Evermann 1896: 459 (type of new genus Zalypnus)


erectus Ginsburg, 1938: 120-121 (type locality northern part of Gulf of California, 26 fathoms; holotype in Bingham Oceanographic Collection)


gulosus (Girard) 1858: 169 (type locality Indianola, Texas). Not Smith, 1936.

Gobius gulosus Girard, 1858:169

Lepidogobius gulosus: Jordan And Gilbert, 1882b: 294 (new combination)[Birdsong 1981]

Microgobius gulosus: Jordan and Eigenmann, 1886:503, 505, 514 (new combination)[Birdsong 1981]


meeki Evermann and Marsh 1900: 356 (type locality between Culebra and Vieques Islands, Puerto Rico at Fish Hawk sta. 6087; holotype USNM 49367)[Birdsong 1981] [Menezes, 1985; Jordan, 1930]

Microgobius omostigma Starks: 1913: 68-69 (type locality Natal, Brazil, in rock pools). Bohlke, 1953: 115 (types in Stanford Univ. collection; SU 22214 designated lectotype, SU 22383 designated paralectotypes).


microlepis Longley and Hildebrand 1940: 257 (type locality Tortugas, Fla.; holotype USNM 108868) [Birdsong 1981]


miraflorensis Gilbert and Starks 1904: 176, fig. 54 (no type locality given) Bohlke, 1953: 115 (SU 6511, Rio Grande, Miraflores, Colombia, given as holotype)


tabogensis Meek and Hildebrand 1928: 873-874, pl. 85 (type locality Taboga Island, Panama; holotype USNM 81844) [Meek and Hildebrand 1928]


thalassinus (Jordan and Gilbert 1882a)

Gobius thalassinus Jordan and Gilbert 1883a: 612 (type locality muddy pools in Charleston Harbor; holotype USNM 29674)[Birdsong 1981]

Microgobius eulepis Eigenmann and Eigenmann 1888(1889): 69-70, holotype MCZ 27123; type locality: Fortress Monroe, South Carolina;  [Birdsong 1981] [Eigenmann and Eigenmann 1888(1889)]

Lepidogobius thalassinus: Jordan and Gilbert, 1883b: 947-948 (new combination; after Jordan and Gilbert, 1883a)[Birdsong 1981]

Microgobius holmesi Smith, 1907: 366-367, fig 168 (type locality Uncle Israel Shoal, Beaufort Harbor, North Carolina)

Microgobius thalassinus: Jordan and Eigenmann, 1886: 503-505 (new combination). Ginsburg, 1934: 35 (M. holmesi and M. eulepis synonymized with M. thalassinus) [Birdsong 1981]


Genus Nes


Nes Ginsburg 1933a: 25-26. Type species: Gobiosoma longum Nichols 1914 by original designation; described as a subgenus of Gobiosoma. [B&R 1968][Nichols 1914]


longus (Nichols 1914)

Gobiosoma longum Nichols 1914: 143, fig. 1 holotype AMNH 5068; Type locality: Florida Keys near Key West. Metzelaar 1919: 139 (specimen from Lagoon at Aruba, Dutch West Indies). Jordan, Evermann and Clark 1930: 446 (compiled). Beebe and Tee-Van 1932: 120 (name only in list of new record of Bermuda fishes); 1933:221 (characters, illustration, Bermuda distribution). Ginsburg 1933b: 26-27 (made type of new subgenus Nes; description, relationships); 1938: 59 (mentioned in connection with elgate first dorsal spine). Longely and Hildebrand 1941: 228 (habits: lives commensally with shrimp; food, color pattern). Hildebrand et al. 1964: 124 (Alacran Reef, Mexico; report on symbiosis with shrimp). [B&R 1968]

Gobiasoma longum: Grant 1927: 45 (genus misspelled; another record of the type specimen disgorged from snapper of doubtful identification). [B&R 1968]

Gobiosoma lividiris Fowler 1944: 140-141, 383, 472 (type locality: Old Providence Island, Caribbean Sea; description based on five small juveniles, characters erroneously recorded). [B&R 1968]


Genus Ophiogobius

This genus was included in the tribe Gobiosominni by Hoese because “it looks like one”. Hense it is included here.  It has eight spines in the first dorsal.


Ophiogobius Gill 1863: 269; Type species: Gobius ophiocephalus Jenyns 1842 = Ophiogobius jenynsi new name, by original designation. [Hoese, 1976:303]


jenynsi new name Hoese 1976

Gobius ophiogobius Jenyns 1842: 97, fig. 3, not of Pallas 1811; Type locality Archipelago de Chonos, Chile. [Hoese, 1976:304]


Genus Palatogobius


 Palatogobius Gilbert, 1971:30 (type species Palatogobius paradoxus Gilbert 1971 by original designation)


paradoxus Gilbert 1971: 31-34, figs. 1,2,3a,4a; holotype ANSP 109182; type locality: St. John, Virgin Islands. [Gilbert 1971]

grandoculus Greenfield 2002:716; holotype USNM 367311; type locality: Cozumel, Mexico.


Genus Pariah


Pariah Bohlke, 1969:2-3, Type species: Pariah scotius Bohlke 1969, by original designation.


Pariah scotius Bohlke 1969: 3-4, fig. 1; holotype ANSP 111855; type locality: sta. TS-52a, Bahamas, Mayaguana Island, SE end of island, 1/2 mi. NW of Southeast Point; 85 ft. 22 Jan. 1968. [Bohlke 1969]


Genus Parrella


Parrella Ginsburg 1938: 116, type species Parrella maxillaris Ginsburg 1938 by original designation. [Ginsburg 1938]


maxillaris Ginsburg 1938: 116-118; holotype in Hancock Coll.; Type locality: La Paz Bay, Gulf of California, 7 fathoms. [Ginsburg 1938]


fusca Ginsburg 1939:57; holotype USNM 107295; Type locality: Miraflores Locks, Panama Canal, lower chamber. [Ginsburg 1939]


ginsburgi Wade 1946: 223; holotype AHF (Allan Hancock Foundation) no. 923; type locality: Balboa, Panama Canal Zone, Republic of Panama, shore collecting. [Wade  1946]


lucretiae (Eigenmann and Eigenmann) 1888(1889): 57-58

Gobius lucretiae Eigenmann and Eigenmann 1888(1889): 57-58; holotype Cal. Acad. of Sci. no. 3957 (number, not location of type, listed in publication); Type locality: Pearl Island, Gulf of Panama. [Eigenmann and Eigenmann 1888(1889)] [Hoese 1983]

                                    Parrella spilopteryx Ginsburg 1939:57; holotype USNM 107293 [Ginsburg 1939][from Hoese, 1975]


macropteryx Ginsburg 1939: 56-57; holotype Bingham Oceanographic Coll. 1688; type locality: Siguanea Bay, Isle of Pines, Cuba. [Ginsburg 1939]


Genus Psilotris


Psilotris  Ginsburg, 1953:22; type species Psilotris alepis, by original designation and monotypy


amblyrhynchus Smith & Baldwin 1999; 433-442; holotype USNM 321019; type locality: Belize, western Caribbean, between Carrie Bow Cay and Twin Cays, lagoon - "shot hole", 15-20 feet.


alepis Ginsburg 1953: 22; holotype USNM 197515; type locality: St. Croix Island, Virgin Islands, on reef, 8 Apr. 1937. [Bohlke 1963] holotype listed by Ginsburg as USNM 123231; type locality: St. Croix Island, Virgin Islands on reef [Ginsburg 1953]


batrachodes Bohlke 1963: 6-7; holotype ANSP 98416; type locality: sta. 304, Bahamas, Sandy Cay (Nassau vicinity), N of the E end, eroded limestone covered with algae, sea-fans, corals, etc., to 20 ft., 17 May 1956.


boehlkei Greenfield 1993: 771-773; holotype ANSP 106059 male; type locality: St. Barthelemy, Les Saintes, Port de Gustavia, southwest of Les Saintes (17o55'N, 62o53'W), 18.3-19.8m, from low coral and sponges on white sand bottom. [Greenfield 1993]


celsus Bohlke 1963: 7-9; holotype ANSP 98427; type locality: sta. 414, coral head N of Green Cay (N of Rose Id.), Nassau vicinity, Bahamas, to 45 ft., 21 July 1957. [Bohlke 1963]


kaufmani Greenfield et.  al.  1993:183-185; holotype: LACM 8939-29, male 32.2mm SL; type locality:Jamaica, Montego Bay, 0.8 km off Chatham Hotel, 15.2 m, in or around rocks [Greenfield et al 1993]


Genus Pycnomma


Pycnomma Rutter 1904: 252, pl. 24 Type species: Pycnomma semisquamatum Rutter 1904, by monotypy; Dichomma used on plates but corrected on explination of plates, p. 254.[B&R 1968][Rutter 1904]


semisquamatum Rutter 1904

Pycnomma semisquamatum Rutter 1904: 252; holotype Dept. of Ichthyology, California Acadamy of Science No. 1478; Type locality: San Jose Island, Gulf of California. Jordan, Evermann and Clark 1930: 438 (compiled; placed in Eleotridae). Ginsburg 1939b: 3 (compared with roosevelti). Fowler 1944: 516 (name only). [B&R 1968][Rutter 1904]


roosevelti Ginsburg 1939b

Pycnomma roosevelti Ginsburg 1939b: 1-3, fig. 1; Holotype USNM 108139; Type locality: Old Providence Island. Schmitt and Schultz 1940: 8 (compiled). Fowler 1944: 471 (name only). [B&R 1968][Ginsburg 1939]


Genus Robinsichtys


Robinsichtys Birdsong, 1988: 438-439; type species R. arrowsmithensis Birdsong 1988, by original designation.


 arrowsmithensis Birdsong 1988: 439-441, figs. 1-3, tables 1-3; holotype USNM 290207; type locality: 21002'N, 86026'W, R/V Gerda sta 952, beginning depth 586 m, ending depth 92 m, 28 Jan. 1968. [Birdsong 1988]


Genus Risor


Risor Ginsburg 1933b: 56 Type species: Garmannia binghami Parr 1930, by original designation [= Garmannia rubra Rosen 1911]; Risor described as a subgenus of Garmannia. [B&R 1968]


ruber (Rosen 1911)

Garmannia hemigymna: Bean 1905: 318 (Green Cay, Bahamas). [B&R 1968]

Garmannia rubra Rosen 1911: 63, fig. 1 Type _____ {type number not listed in publication}; Type locality: "among sponges, corals etc. near the shore at Mastic Point, Andros," Bahamas. Metzelaar 1922: 139 ("Many specimens from sponges in Caracasbay... and Spanish water" Curacao). [B&R 1968] [Rosen 1911]

Garmannia (Risor) rubra: Ginsburg 1933b: 59 (synopsis after Rosen; comparison with binghami). [B&R 1968]

Risor ruber: ginsburg 1939a: 61 ( compared with mirus). Longley and Hildebrand 1941: 228 (Tortugas, Floida; binghami placed in the synonymy of ruber). [B&R 1968]

Risor rubra: Fowler 1944: 450 (in check-list of Bahama fishes). [B&R 1968]

Garmannia (Risor) binghami: Ginsburg 1933b: 58 (redescription of type specimen) [B&R 1968]

Risor binghami: Ginsburg 1939a: 61 (compared with mirus). Pearse 1950: 150 (Bimini, Bahamas; eight speciemens removed from the sponge, Hiricina strobilina (Lamarck), 1-2 m depth). [B&R 1968]

Risor mirus Ginsburg 1939a: 60; holotype Bingham Oceanographic Coll. 1691; (Type locality: Siguanea Bay, Isle of Pines, Cuba). [Ginsburg 1939][B&R 1968]

?Microgobius spec.: Metzelaar 1919: 138 (a sepcimen from "Bay of Asiento, Curacao" was placed in the synonymy of mirus by Longley and Hildebrand (1941: 228)). [B&R 1968]



Genus Varicus


Varicus Robins and Bohlke, 1961: 46; Type species: Varicus bucca Robins and Bohlke 1961:47 by original designation.


bucca Robins and Bohlke 1961: 47-49; holotype ANSP 93083; Type locality: Saba Bank between St. Kitts, British West Indies and St. Croix, Virgin Islands, Lat 17033'N., Long. 63035'W., 125-132 fathoms, M/V Oregon, Station 2356. [R&B 1961]


imswe Greenfield 1981: 269-271; holotype FMNH 83894; type locality: Carrie-Bow Cay, Belize, at the barrier-reef dropoff east of cay, depth 21-25 m. [Greenfield 1981]


marilynae Gilmore 1979: 126-128; holotype USNM 218406; Type locality: Atlantic Ocean off Sebastian Inlet, Brevard and Indian River counties, Florida; 27050.3'N, 79057.0'W; 61-79 m [Gilmore 1979]







Acero, P.A. 1981. Two new species of gobiid fishes from the Colombian Pacific. Jap. J. Ichthy. 28(3): 243-246.


Beebe, W. & G. Hollister. 1933. New species of fish from the West Indies. Zoologica 12(9): 82-88. 3 fig. (Scientific Cont. of the NY Zool. Soc.)(NOTE: Vol. 12 includes articles from April 30, 1929 to Aug. 30, 1934; the publication date of the vol. is listed as 1931 in Bioabstracts]


Beebe, W.; J. Tee-Van. 1928. The fishes of Port-au-Prince Bay, Haiti, with a summary of the known species of marine fish of the island of Haiti and Santo Domingo. Zoologica, N.Y., 10(1): 1-279.


Birdsong, R. S. 1967. The goby named Microgobius cinctus Nichols, a junior synonym of Lythrypus dalli (Glibert). Copeia 1967: 466-467.


_________. 1968. Microgobius crocatus, a new gibiid fish from Pacific Panama. Proc. Biol. Soc. Wash. 81:191-196.


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Birdsong, R. S. 1981. A review of the gobiid fish genus Microgobius Poey. Bull. mar. Sci. 31(2):267-306.


Birdsong, R. S. 1988. Robinsichthys arrowsmithensis, a new Genus and species of deep-dwelling Gobiid fish from the Western Carribean. Proc. Biol. Soc. Wash. 101(2): 438-443.


Birdsong, R.S. and C.R. Robins.  1995.  New genus and species of seven-spined goby (Gobiidae: Gobiosominni) from the offing of the Amazon River, Brazil.  Copeia 1995:676-683.


Bleeker, P. 1874. Esquisse d'un système naturel des Gobioides. Arch. Neerland. Sci. Nat., 9: 289-331.


Bohlke, J.E. 1953. A catalogue of the type specimens of recent fishes in the natural history museum of Stanford University. Stanford Ichthy. Bull. 19: 135-186.


Bohlke, J.E. 1963. The species of the west Atlantic Gobioid fish genus Psilotris. Notulae Naturae No. 362: 1-10.


Bohlke, J. E. 1969. Pariah scotius, A new sponge-dwelling gobiid fish from the Bahamas. Notulae Naturae No. 421: 1-7.


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Bohlke, J.E. and C.R. Robins. 1960. Western Atlantic gobies of the genus Lythrypnus, with notes on Quisquilius hipoliti and Garmannia pallens. Proc. Acad. Nat. Sci. Phila., 112: 73-98, 2 figs., 3 pls.


Bohlke, J.E. and C. R. Robins. 1968. Western Atlantic seven-spined gobies, with description of ten new species and a new genus, and comments on Pacific relatives. Proc. Acad. Nat. Sci. Phil., 120(3) 45-174.


Bohlke, J. E. and C. R. Robins. 1969. Western Atlantic sponge-dwelling gobies of the genus Evermannichthys: Their taxonomy, habits and relationships. Proc. Acad. Nat. Sci. Phil. 121(1): 1-24.


Breder, C.M., Jr. 1927. The fishes of the Rio Chucunaque drainage, eastern Panama. Bull. Am. Mus. Nat. Hist. 57: 91-176.


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Chirichigno, F.N. 1968. Nuevos registros para la ictiofauna marina del Peru. Inst. del Mar del Peru. Peru Boletin 1: 379-503.


Chirichigno, F.,N. 1978. Nuevas adiciones a la ictiofauna marina del Peru. Informe Inst. Mar. Peru No. 46: 1-109.


Colin. P.L. 1974. Comparative biology of the gobies of the genus Gobiosoma subgenus Elacatinus in the western north Atlantic Ocean. Dissertation Abstr. int. (B) 35(12):339


Colin, P. 1975. The neon gobies. The comparative biology of the gobies of the genus Gobiosoma, subgenus Elacatinus, (Pisces: Gobiidae) in the tropical western North Atlantic Ocean. T.F.H. pub., Neptune City, NJ 1-304.


Crabtree, R.E. and D.P. Middaugh. 1982. Oyster shell size and the selection of spawning sites by Chasmodes bosquianus, Hypleurochilus geminatus, Hypsoblennius ionthas (Pisces: Blenniidae) and Gobiosoma bosi (Pisces: Gobiidae) in two South Carolina estuaries. Estuaries 5(2): 150-155.


Dahlberg, D., Conyers, J.C. 1973. An Ecological study of Gobiosoma bosci and G. ginsburgi (Pisces, Gobiidae) on the Georgia Coast. Fishery Bulletin 71(1):279-287.


Dawson, C.E. 1963. Notes on Gobiosoma longipala Ginsburg, with a record of its occurence in Mississippi waters. Copeia No. 3: 585-586.


Dawson, C.E. 1966. Studies on the Gobies (Pisces: Gobiidae) of the Mississippi Sound and Adjacent Waters. I. Gobiosoma. Amer. Midl. Natur. 76(2): 379-409.


Dawson, C.E. 1968. An Adventitious Dorsal Fin-Ray in Microgobius gulosus (Pisces: Gobiidae). Anat. A_z. Bd. 122: 498-501.


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Dawson, C.E. 1969. New Seven-spined Goby, Gobiosoma (Austrogobius) polyporosum, from the Pacific Coast of Panama. Copeia 1969 No. 3: 510-514.


Dawson, C.E. 1971. Gobiosoma (Garmannia) yucatanum, a new seven-spined Atlantic Goby from Mexico. Copeia No.3: 432-439.


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de Buen, F. 1951. Contribuciones a la Ictiologia. V-VI. Sobre algunas especies de Gobiidae de la collecion del Laboratoire Arago (Banyuls-Sur-Mer, Francia) y descripcion de un nuevo genero (Austrogobius) sudamerica-no. Bol. Inst. Paulista Oceanogr., 2(2): 55-73, 1 pl.


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Eschmeyer, W.N. 1990. Catalog of the Genera of Recent Fishes. Calif. Acad. of Sci. Pub. 697pp.


Evermann, B.W., and B.A. Bean 1898. Indian River and its fishes. U.S. Fish. Comm. Rep. 22(1896): 227-262.


Evermann and Marsh, 1899. Rept. U.S. Fish Comm., XXV, {Reference should read 1900} [Jordan, 1930]


Evermann, B.W., and M.C. Marsh 1900. Description of new genera and species of fishes from Puerto Rico. U.S. Comm. Fish and Fish. Rep. 25(1898/1899): 351-362.


__________. 1902. The fishes of Porto Rico. Investigations of the aquatic resources and fisheries of Porto Rico by the U.S. Fish Commission Steamer Fish Hawk in 1899. Bull. U.S. Fish Comm. 20:49-350.


Feddern, H.A. 1967. Larval development of the neon goby Elacatinus oceanops , in Florida. Bul. of Marine Sci. 17(2): 367-375.


Findley, L.T. 1975. A new species of goby from Malpelo Island (Teleostei: Chriolepis). Smithsonian Contr. Zool. 176: 94-98.


Fischer, W. (ed.), 1978. FAO species identification sheets for fishery purposes. Western Central Atlantic (fishing area 31). Vols. 1-7. Roma, FAO, pag. var.


Fowler,H. W. 1931. Fishes obtained by the Barber Asphalt Company in Trinidad and Venezuela in 1930. Proc. Acad. Nat. Sci. Philadelphia, 83: 391-410.


Fowler, H.W. 1941. Notes on Florida fishes with descriptions of seven new species. Proc. Acad. Nat. Sci. Phila. 93: 81-106, 17 figs.


Fowler, H. W. 1945. A study of the fishes of the southern piedmont and coastal plain. Acad. Nat. Sci. Phila. Monograph 7: 401 pp.


Fowler, H.W. 1949. Description of a new species of goby (Gobiosoma thomasi) from Jamaica. Notulae Naturae No. 218: 1-3. {note: =Sicydium plumieri (Bloch) in Erdman (1961:450)}


Fowler, H. W. 1950. Results of the Catherwood-Chaplin West Indies Expedition, 1948. Part III. The Fishes. Proc. Acad. Nat. Sci. Philadelphia, 102: 69-93.


Freihofer, W.C. 1970. Some nerve patterns and their systematic significance in paracanthopterygian, salmoniform, gobioid, and apogonid fishes. Proc. Calif. Acad. Sci. 38:215-263.


Gilbert, C.H. 1891. Scientific results of explorations by the U.S. Fish Commission Steamer Albatross. No. XII - a preliminary report on the fishes collected by the steamer Albatross on the Pacific coast of North America during the year 1889, with descriptions of twelve new genera and ninety-two new species. Proc. U.S. Nat. Mus. 13(1890): 49-162.[NOTE: vol. 13(1890) was published in 1891]


Gilbert, C.H. 1892. Scientific results of explorations by the U.S. Fish Commission steamer ALBATROSS, XXII: Description of thirty-four new species of fishes collected in 1888 and 1889, principally among the Santa Barbara Islands and in the Gulf of California. Proc. U.S. Natl. Mus. 14(880): 539-566. [NOTE: Vol. 14(1891) was published in 1982]


Gilbert, C. R. 1971. Two new genera and species of western Atlantic gobiid fishes with vomerine teeth. Copeia 1971 (1):27-38.


Gilbert, C.R. and G.H. Burgess. 1986. Variation in Western Atlantic Gobiid Fishes of the Genus Evermannichthys. Copeia 1986(1): 157-165.


Gilbert and Starks. 1904. The fishes of Panama Bay. Mem. Calif. Acad. Sci., 4: 1-304.


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Gilmore, G.R. 1979. Varicus marilynae, a new Gobiid Fish from Florida. Copeia 1979(1): 126-128.


Ginsburg, I. 1933a. Description of new and imperfectly known species and genera of gobioid and pleuronectid fishes in the United States National Museum. Proc. U. S. Nat. Mus., 82(2961): 1-23.


Ginsburg, I. 1933b. A revision of the genus Gobiosoma (family Gobiidae) with an account of the genus Garmannia. Bull. Bingham Oceanogr. Coll., 4(5): 1-59.


Ginsburg, I. 1934. The distinguishing characters of two common species of Microgobius from the east coast of the United States. Copeia 1934: 35-39.


Ginsburg, I. 1935. Bollmania litura, a new species of Goby. Smith. Misc. Coll. Vol. 91(20): 1-3.


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Ginsburg, I. 1938b. Eight new species of gobioid fishes from the American Pacific coast. Allan Hancock Pacific Expedition 2: 109-121.


Ginsburg, I. 1939a. Twenty one new American gobies. J. of the Washington Academy of Sciences; 29(2):51-63.


Ginsburg, I. 1939b. Two new gobioid fishes collected on the presidential cruse of 1938. Smithsonian Misc. Colls., 98(14): 1-5.


Ginsburg, I. 1942. Seven new American fishes. Journ. Washington Acad. Sci., 32(12): 364-370.


Ginsburg, I. 1944. A description of a new gobiid fish from Venezuela, with notes on the genus Garmannia. Journ. Washington Acad. Sci., 34(11): 375-380.


Ginsburg, I. 1953. Ten new American gobioid fishes in the United States National Museum, including additions to a revision of Gobionellus. Journ. Washington Acad. Sci., 43(1): 18-26.


Girard, C. 1858. Notes upon various new genera and new species of fishes, in the museum of the Smithsonian Institution, and collected in connection with the United States and Mexican Boundary Survey: Major William Emory, Commissioner. Proc. Acad. Nat. Sci. Philadelphia, 1858: 167-171.


Greenfield, D.W. 1981. Varicus imswe, a new species of gobiid fish from Belize. Copeia 1981(2): 269-272.


Greenfield, D.W. 1993.  New Goby, Psilotris boehlkei (Pisces: Gobiidae), from the Western Atlantic, with a key to the species.  Copeia 1993(3): 771-775.


Greenfield, D.W., Findley, L.T. and R.K. Johnson.  Psilotris kaufmani n.  sp.  (Pisces: Gobiidae), a Fourth Western Atlantic Species of Psilotris.  Copeia 1993(1): 183-186.


Gánther, A. C. L. G. 1861a. On a collection of fishes sent by Capt. Dow from the Pacific coast of Central America. Proc. Zool. Soc. London, 1861: 370-376.


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Hildebrand, S.F., and L.E. Cable. 1938 Further notes on the development and life history of some teleosts at Beauford, N.C. Bull. Bur. Fish. 48: 504-642.


Hildebrand, S.F. and W.C Schroeder. 1928. Fishes of the Chesapeake Bay. Bull. U.S. Bur. Fisheries 43(1) [Doc. 1024]: 1-366, 211 fig., 1 map 1927(1928).


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Hoese, D. F. 1978. Eleotridae in W. Fischer (ed.)


Hoese, D. F. 1978. Gobiidae, in W. Fischer (ed.)


Hoese, D.F. 1985. Revision of the Eastern Pacific Species of the Genus Barbulifer (Pisces: Gobiidae). Copeia 1985(2): 333-339.


Hubbs, C.L. 1921. Description of a new Genus and species of Goby from California with notes on related species. Occasional Papers of the Mus. of Zool., Univ. Mich. No. 99: 1-4.


Hubbs, C.L. 1923. A note on the species of Evermannichthys, a genus of sponge-inhabiting gobies. Occasional papers of the Mus. of Zool. No. 144: 1-2.


Hubbs, Carl L. 1960. The marine vertebrates of the outer coast. In Symposium: the biogeography of Baja California and adjacent seas. Part II. Marine Biotas. Syst. Zool. 9:134-147.


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Jordan, D. S. 1904. Notes on fishes collected in the Tortugas Archipelago. Bull. U. S. Fish Comm., 22 for 1902: 539-544.


Jordan, D.S. and C.H. Bollman. 1890. Scientific Results of Exploration by the U.S. Fish Commission Steamer Albatross. No. IV.- Descriptions of new species of fishes collected at the Galapagos Islands and along the coast of the United States of Colombia, 1887-'88. Proc. of the U.S. Nat. Museum. Vol XII(770)(1889): 149- ???


Jordan, D.S. and C.H. Eigenmann. 1886. A review of the gobiidae of North America. Proc. U.S. Nat. Mus. 9:477-518.


Jordan, D.S. and B.W. Evermann. 1896. A checklist of fishes and fish-like vertebrates of North and Middle America. U.S. Comm. Fish. Fish Rep. 21(1895):207-584.


Jordan, D. S.; B. W. Evermann. 1898. The fishes of North and Middle America: ... Bull. U. S. Nat. Mus., 47(3): 2183-3136.


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Jordan, D. S.; C. H. Gilbert. 1882a. Description of nineteen new species of fishes from the Bay of Panama. Bull. U.S. Fish. Comm. 1:306-335.


Jordan, D.S. and C.H. Gilbert. 1882b. Notes on fishes observed about Pensacola, Florida, and Galveston, Texas, with description of new species. Proc. U.S. Nat. Mus. 5:241-307.


Jordan, D.S. and C.H. Gilbert. 1882c. List of fishes collected at Panama by Charles H. Gilbert. Bull. U.S. Fish. Comm. 2(1882): 109-111.


Jordan, D.S. and C.H. Gilbert. 1883a. Notes on a collection of fishes from Charlston, South Carolina, with descriptions of three new species. Porc. U.S. Nat. Mus. 5: 580-620.


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Jordan, D. S.; C.H. Gilbert. 1884. Descriptions of ten new species of fishes from Key West, Florida. Proc. U. S. Nat. Mus., 7(402): 24-32.


Jordan, D.S. and E.C. Starks. 1895 Fishes of Sinaloa. Proc. Calif. Acad. Sci., 2d ser., V: 1-498.


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Koumans, F.P. 1931. A preliminary revision of the genera of the gobioid fishes with united ventral fins. Drukkerij "Imperator" N.V., Lisse (Curacao): 1-174.


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Longley, W.H. and S.F. Hildebrand. 1940. New genera and species of fishes from Tortugas, Florida. Papers from the Tortugas Laboratory, Vol. 32. Carnegie Instit. Wash. Publ. 517: 223-285.


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Nichols, J.T. 1952. Four new gobies from the eastern and western Pacific. Am. Mus. Novitates (1954): 5pp.


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Richardson, S.L., and E.B. Joseph. 1975. Occurance of larve of the green goby, Microgobius thalassinus, in the York River, Virginia. Ches. Sci. 16: 215-218.


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Robins, C. R. 1960. Garmannia saucra, a new gobiid fish from Jamaica. Proc. Biol. Soc. Washinton, 73: 281-285.


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Robins, C. R.; J. E. Bohlke. 1964. Two new Bahaman gobiid fishes of the genera Lythrypnus and Garmannia. Notulae Naturae, no. 364: 1-6.


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