The
American Seven-spined Gobies
Tribe Gobiosomatini
Species
Documentation
Prepared by:
Research Associate in Residence
Dept. of Biology
114 Hofstra University
Hempstead, NY 11549
Revision Date: Aug. 29, 2002
This is a working document
and not a completed list of all references, synonoyms, etc. If you see any mistakes or wish to contribute
additional information, please e-mail
me.
The following genera are included in alphabetical order:
Aboma, Akko, Aruma, Barbulifer, Bollmannia
, Chriolepis, Eleotrica , Enypnias, Evermannichthys, Ginsburgellus, Gobiosoma
(including the subgenera – Austrogobius, Garmannia , Gobiolepis,
Gobiosoma, Elacatinus, Tigrigobius ), Gobulus, Gymneleotris,
Microgobius, Nes, Ophiogobius, Palatogobius, Pariah, Parrella, Psilotris,
Pycnomma, Robinsichtys, Risor, Varicus.
KEY: Information within the
brackets [ ] indicates the source of the information and that the information
has been checked against the original publication.
NOTE: The classification used for Gobiosoma
is that of Böhlke & Robins 1968,
when ever possible. While both Hoese
(1971) and Van Tassell (1998) have proposed new classifications for all or
part of the group in their Ph.D. dissertations, neither is currently published. When I have sufficient information to resolve
a few remaining nodes, I’ll publish a new phylogeny and changes will then
be made to this page.
Genus Aboma
Aboma Jordan and Starks
1895; Type species Aboma etheostoma Jordan and Starkes; Fishes of Sinaola,
Proc. Calif. Acad. Sci 2d ser., 1895: 497. [Jordan, 1930]
etheostoma Jordan
and Starks 1895; locality: Mazatlan, Mexico.
polyporosum Dawson
1969: 510-513; holotype USNM 203573; type locality: Panama, Pacific coast
from deep tidepool at Farfan Point.
Genus Akko
Akko Birdsong and Robbins 1995
dionaea Birdsong &
Robins 1995: 676, by original designation; holotype USNM 329524; type locality:
Amapa Prov. East of Iiha de Maraca, Brazil.
Genus Aruma
Aruma Ginsburg 1933a:
16 Type species: Gobiosoma occidentale Ginsburg 1933:16-17, by original
designation; erected as a subgenus of Gobiosoma.[Gin. 1933a]
histrio (Jordan 1884)
Gobiosoma
histrio Jordan 1884:260; holotype USNM 35147; Type locality: Guaymas,
Mexico [Hoese 1976]{Proc. U.S. Nat. Mus. for 1884 was published in 1885}
Gobiosoma occidentale
Ginsburg 1933a; Type USNM 92801; Type locality: La Paz Harbor, Mexico;[Gin.
1933] [syn. by Hoese 1976 Copeia No. 2]
Genus Barbulifer
Barbulifer Eigenmann
and Eigenmann 1888(1889): 70 Type species: Barbulifer papillosus Eigenmenn
and Eigenmann 1888(1889): 70 [= Gobiosoma ceuthoecum Jordan and Gilbert
1884] by original designation, see Böhlke & Robins 1968: 120 for
additional information on problems associated with the type species. [B&R
1968] [Eigenmann and Eigenmann 1888(1889)][Hoese 1985]
ceuthoecus (Jordan
and Gilbert 1884)
Gobiosoma ceuthoecum
Jordan and Gilbert 1884: 29, holotype USNM 34970; Type locality: Key West,
Florida, from the cavity of a sponge. Jordan 1887: 894 [106] (in check=list).
Jordan and Eigenmann 1887: 506, 508 (synopsis of species).[B&R 1968] [Jordan
and Gilbert 1884]
Barbulifer ceuthoecus:
Eigenmann and Eiganmann 1888: 72 (placed in Barbulifer, with papillosus).
Jordan and Evermann 1896: 461 (in check-list). Jordan and Evermann 1898: 2260
(description). Robins 1958: 196 (habits in Florida Keys). Hildebrand, Chavez
and Compton 1964: 123 (Alacran Reef, Yucatan). [B&R 1968]
Barbulifer cauthoecus:
Jordan, Evermann and Clark 1930: 446 (species name misspelled in check-list).
Erdman 1956: 336 (Guanica, Puerto Rico; we did not examine the collection
from Guajataca). [B&R 1968]
Barbulifer papillosus
Eigenmann and Eiganmann 1888(1889): 70-72; holotype 26186 MCZ; type locality:
Key West Florida; distinguished from ceuthoecus). [B&R 1968] [Eigenmann
and Eigenmann 1888(1889)]
Barbulifer sp.: Caldwell
and Caldwell 1964: 29 (Caledonia Bay, Panama).[ B&R 1968]
Barbulifer, species
(in part): Caldwell 1966: 69 (Jamaica collection listed, some of them this
species, others antennatus).[B&R 1968]
antennatus Böhlke
and Robins 1968
Barbulifer antennatus
Böhlke and Robins 1968: 126; Holotype ANSP 110642; Type locality: sta. 410,
Bahamas, Grand Bahamas Bank, Exuma Cays, baylet on SE side of Warderick Wells
Cay; to ca. 6 ft. [B&R 1968]
Barbulifer, species
(in part): Caldwell 1966: 69 (some Jamaican collection listed are of this
species, others are ceuthoecus). [B&R 1968]
mexicanus Hoese 1985
Barbulifer mexicanus Hoese
1985: 337-338; holotype SIO 61-248; type locality: Los Frailes, Baja California,
Gulf of Mexico. [Hoese 1985]
pantherinus (Pellegrin
1901)
Gobiosoma pantherinum
Pellegrin 1901: 165; syntypes MNHN 01-295; Type locality: Sargassum floating
in the Gulf of California. Ginsburg 1933b: 17 (statement that pantherinum
probably belongs to Barbulifer). [B&R 1968] [Hoese 1985- only
source to list the type no.][Pellegrin 1901]
Gobiosoma spiriti-sancti
Wade 1946: 219, pl. 30; holotype LACM 21556; type locality: Espiritu Santo
Island, Gulf of California, Mexico). [B&R 1968] [Hoese 1985]
Genus Bollmannia
Bollmannia Jordan,
1890: 164(in: Jordan and Bollman 1890); type species Bollmannia chlamydes
Jordan 1890: 164-165, by original designation.
chlamydes Jordan 1890:164-165;
Lectotype USNM 93825, by subsequent designation of Ginsburg 1939: 58; Type
locality: Panama Bay; Albatross station 2800, lat. 8051', long.
79031'30"; 7 fathoms. Note: Jordans original type, USNM 41158,
does not match the description of the type and the jar contain two distinct
species. [Ginsburg 1939]
boqueronensis Evermann
and Marsh 1900: 356; holotype USNM 49366; type locality: sta. 6074 off Puerto
Real, Puerto Rico, dredged by the Fish Hawk, in 8.5 fathoms of water.
[Evermann and Marsh 1890]
communis Ginsburg
1942:364-365; holotype USNM 119873; type locality: Pelican station
77-1; 28059'N., 89029'W., 10 fathoms. [Ginsburg 1942]
eigenmanni Gamon
Gobius eigenmanni
Gamon ??? USNM 231318
jeannae Fowler 1941:
95-96; holotype ANSP 69718; type locality: Key West, Florida.
litura Ginsburg 1935:
1-3; holotype USNM 93797; type locality: Caroline station 59. Samana Bay,
Dominican Republic, 18-19 fathoms. [Ginsburg 1935]
longipinnis Ginsburg
1939: 60; holotype Bingham Oceanographic Collection 1690; Type locality: Angeles
Bay, Gulf of California. [Ginsburg 1939]
macropoma Gilbert
1892:556; syntypes (6) SU 81 [Böhlke 1953]; type locality: Gulf of California
just north of La Paz Bay, at Sta. 2996 (Albatross Exp.); 112 fathoms. [Gilbert
1892]
marginalis Ginsburg
1939: 59; holotype USNM 107284; type locality: Solango Island, Ecuador, 12
fathoms. [Ginsburg 1939]
ocellata Gilbert 1892:557;
type not designated by Gilbert; Type locality: Northern part of the Gulf of
California; Albatross Exp. Sta. 3031 and 3035, 30 and 33 fathoms. [Gilbert
1892] Lectotype USNM 107286 by subsequent designation of Ginsburg 1939: 59-60.
Gilbert's original specimens were lost. His discription is based on a combination
of three species. The only jar at the USNM labeled ocellata from the
type locality was used to designate the lectotype. However, since it is not
certain if Gilbert ever examined these specimens it may be a neotype and not
a lectotype.
pawneea Ginsburg 1939:
60; holotype Bingham Oceanographic Collection 1689; Type locality: Perlas
Islands, Panama Bay, lat.8029'40", long. 78052'30",
19-24 fathoms. [Ginsburg 1939]
stigmatura Gilbert
1892: 556; syntype SU 10 [Böhlke 1953]type USNM_____; type locality: northern
part of the Gulf of California, Albatross Exp. Sta. 3016 and 3017, 76 and
58 fathoms [Glibert 1892]
umbrosa Ginsburg 1939:
59; holotype USNM 107290; type locality: Panama Bay, Albatross station 2804,
lat. 8016'30", long. 79037'45", 47 fathoms.
[Ginsburg 1939]
Genus Chriolepis
Chriolepis Gilbert,
1892:557;Type species Chriolepis minutillus Gilbert 1892:558 by original
designation. [Gilbert 1892]
minutillus Gilbert 1892:
558; holotype USNM 48261 [Gilmore 1979] type locality: Albatross Exp.
Sta. 2825, Gulf of California [Findley 1975] [Gilbert 1892]
cuneata Bussing 1990:113-115.
(type locality: Isla San Jose, Islas Murcielagos, Costa Rica) depth 10-14
meters. holotype LACM 4428-1; paratypes also at LACM
dialepta Bussing 1990:115-117.
(type locality: East side Isla del Coco, 300 m SW Cabo Desubierta, in 20 meters
depth); holotype LACM 44821-4; paratypes also at LACM
Chriolepis walkeri,
Findley 1983, unpublished Ph.D. dissertation, Univ. of Arizona, 193p.
fisheri Herre 1942:
297; holotype _____ not listed in publication; type locality: Barbados, depth
45 fathoms. [Herre 1942]
lepidotus Findley
1975:94-97; holotype USNM 211456; type locality: Malpelo Island, Colombia,
east side of island, depth ca. 10 meters, above the coral zone, cobble bottom
with some calcareous sand and boulders, very sparse algae growth. [Findley
1975]
tagus Ginsburg 1953:21-22;
holotype USNM 123232; type locality: Tagus Cove, Albemarle Island, Galapagos
Archipelago, 10-18 fathoms. [Ginsburg 1953]
vespa (Hastings &
Bortone) 1981: 428-432; holotype USNM 221523; type locality: northeastern
Gulf of Mexico, 28024'N, 85015'W, R/V Bellows (cruise
Bureau of Land Management, BLM 15, station III-C, 183 m, 26 July 1975. [H&B
1981]
Eleotriculus (subgenus) Ginsburg 1938: 111; type species Chriolepis zebra Ginsburg 1938 by original designation. [Ginsburg 1938]
zebra Ginsburg 1938:
109-111; holotype in Hancock Coll.; Type locality: San Gabriel Bay, Espiritu
Santo Island, Gulf of California, shore collection. [Ginsburg 1938]
Genus Eleotrica
Eleotrica Ginsburg
1933a: 10; Type species: Eleotrica cableae Ginsburg 1933, by original
designation.
Alepidoeleotris Herre 1935:413; synonym of Eleotrica Birdsong and Robins 1995:682
cableae Ginsburg 1933a:11-12;
type USNM 65517; Type locality: Chatham Island, Galapagos Archipelago [G.
1933]
Alepidoeleotris tigris
Herre (1935:413) synonomized by Myers (1936:129) without checking the specimen;
Eschmeyer and Bailey (1990:19) without discussion; Birdsong and Robins (1995:682)
with discussion.
Genus Enypnias
Enypnias Jordan and
Evermann 1898: 2233-2234. Type species: Gobius seminudus Gunther 1861,
by original designation; proposed as a subgenus of Garmannia. [B&R
1968]
seminudus Günther)
1861a
Gobius seminudus Günther
1861:a 372; Type ____ (number and location of specimen not listed in original
publication); Type locality: west coast of Central America. 1861b: 554 (description);
1862:328 (repeat of original description). [B&R 1968] [Günther 1861a]
aceras Ginsburg 1939a:
61; Holotype USNM 107298; Type locality: Miraflores Locks, lower chamber,
Pacific slope, Panama Canal. [Ginsburg 1939a][B&R 1968]
Enypinas aceras: Hildebrand
1939: 43 (genus misspelled; 127 specimens recorded from upper and lower chambers
of Miraflores Locks). [B&R 1968]
Genus Evermannichthys
Evermannichthys Metzelaar,
1919: 139-140 figs 39,40. Type species: Evermannichthys metzelaari
Hubbs 1923:2 (by monotypy).
Radcliffella Hubbs,
1921: 2; Type species Garmannia spongicola Radcliffe 1917, by original
designation. [B&R 1969]
spongicola (Radcliffe
1917)
Garmannia spongicola Radcliffe,
1917: 423-425, fig. (unnumbered); Holotype USNM 77501; Type locality: North
Carolina, 1 mile SE of fishing buoy, 34019'N, 76048'
W in 14: fms., from
sponge. Hubbs, 1921: 2 (made type of new genus Radcliffella). Kousmans,
1931: 72-73 (characters, nomenclature; retained in Evermannichthys).
[B&R 1969][Radcliffe 1917]
Evermannichthys spongicola:
Hubbs, 1923: 1-2 ( spongicola placed in Evermannichthys Metzelaar
and compared to E. metzelaari). Hildebrand, 1941: 229 (North Carolina;
compiled). Delsman, 1941: 76 (9 specimens from near Tampa). Springer, 1961:
482 (records near Tampa Bay, compiled). Moe et al., 1966: 75 (4 localities
on West Coast of Florida south of Tampa Bay). [B&R 1969]
Evermannichthys metzelaari
(misidentifications): Pearse, 1932: 119 (Tortugas; reported from sponge,
Speciospongia vespara [sic]). Longley and Hildebrand, 1941: 229-230
(in part; descriptive data on 3 specimens from Tortugas). Pearse, 1950: 151
(taken at Tortugas; compiled). Briggs, 1958: 288 (in part; Florida distribution).
Duarte-Bello, 1959: 122 (reference to eastern Florida only). Bullis and Thompson,
1965: 56 (Gulf of Mexico records). [B&R 1969]
bicolor Thacker 2001
Evermannichthys bicolor Thacker 2001:2-4, fig 1; Holotype LACM 54100-3; Type locality: Navassa Island, Caribbean Sea, rotenone station in a flat sand habitat with scattered rocks, gorgonians, and sponges at a depth of 27-30 feet.
convictor Böhlke
and Robins 1969
Evermannichthys convictor
Böhlke and Robins 1969: 11-13, fig. 2;by original designation; Holotype
ANSP 111862; Type locality: sta. TS-7, Bahamas, Graet Bahama Bank, Green Cay
(E of southern Andors ISland), W side of cay, 100 yards S of sand spit at
far NW end; 45 ft.; 10 Jan. 1968. [B&R 1969]
metzelaari Hubbs 1923
Evermannichthys spongicola
Metzelaar, 1919: 139-140, figs 39-40 (description; type locality; Curacao);
1922: 141 (generic diagnosis). [B&R 1969]
Evermannichthys metzelaari
Hubs, 1923:1-2 (substitute name for E. spongicola, preoccupied as result
of referral to Evermannichthys of Garmannia spongicola Radcliffe).
Beebe and Tee-Van, 1928: 223-224 (characters, ecology, Haitian records). Koumans,
1931: 72-73 (nomenclature, discussed under the names Evermannichthys spongicola
and E. metzelaari plus two, never used, manuscript manes Evermannella
spongicola and Diogenides cavitarius). Longley and Hildebrand,
1941: 229-230 (in part; references to Dutch West Indies only). Briggs, 1958:
288 (in part; Lessrer Antilles only). Duarte-Bello, 1959: 122 (in part, Lesser
Antilles reference only). Böhlke and Chaplin 1968:627 (characters, Bahaman
record, distribution, figure). [B&R 1969]
silus Böhlke
and Robins 1969
Evermannichthys silus
Böhlke and Robins 1969: 10-11,fig. 1; by original designation; Holotype
ANSP 111865; Type locality: sta. TS-71, Bahamas, Samana (Atwood) Cay, E of
the Graet Bahama Bank, middle of W half of S side of cay; 90 ft. 27 Jan. 1968.
[B&R 1969]
Genus Ginsburgellus
Ginsburgellus Böhlke
and Robins 1968: 140-141. Type species Gobiosoma novemlineatum Fowler
1950 = Ginsburgellus novemlineatus, the only species in the genus.
[B&R 1968]
novemlineatus (Fowler
1950)
Gobiosoma novemlineatum
Fowler 1950: 89, figs. 44-46. holotype ANSP 71778; Type locality: Caribbean
Sea, San Andres [St. Andrews Island] off coast of Nicaragua. Erdman 1956: 335 (Puerto Rico; color; association
with sea urchin, Echinometra). [B&R 1968][Bohlke, E.B. 1984, for holotype
number]
Genus Gobiosoma
Gobiosoma Girard 1858:
169 (type species: Gobiosoma molestum Girard 1858 = Gobiosoma bosc
Lacepède 1800, by subsequent designation by Bleeker, 1874: 310)
Subgenus Gobiosoma
Gobiosoma Girard 1858:
169 (type species: Gobiosoma molestum Girard 1858 = Gobiosoma bosc
Lacepède 1800, by subsequent designation by Bleeker, 1874:310)
Dilepidion Ginsburg 1933a: 17; Type species: Gobiosoma
ginsburgi Hildebrand and Schroeder 1928, by original designation.[Ginsburg
1933a]
bosc (Lacepède, 1798)
Gobius bosc Lacepède
1800:555 (Charlston, S.C., from a manuscript by Bosc.; as restricted by Hildebrand
and Schroeder.)[Gins. 1933b]
Gobius alepidotus
Bloch and Schneider 1801 (Apparently based on Lacepède's account)[Gins. 1933b]
Gobius viridipallidus
Mitchil 1815 (New York; as restricted by Cuvier and Valenciennes)[Gins.
1933b]
Gobiosoma molestum
Girard 1858 (Texas; type reexamined; based on a specimen of the present species).
Girard; 1859(Texas, based on same specimen as preceding record). Günther;
1861 (Texas, after Girard). Jordan and Gilbert, 1882 (Texas after Girard).[Gins.
1933b]
Gobiosoma alepidota
Uhler and Lugger; 1876 (Worcester County, Maryland). [Gins. 1933b]
Gobiosoma bosci Bean
1889 (Somers Point and Ocean City, N.J.). Bean 1890(Long Island, New York).
Bean, 1892 (Cape Charles, Va.). Evermann
and Kendall, 1894 (Galveston and Corpus Christi, Texas). Smith and Bean (in
part), 1898 (Potomic River at Gunston Wharf, VA). Evermann, 1898, 1899 {could
be 1900 ref. see bib.}, (Baldwin Lodge, Miss.). Bean, 1898 (Long Island, many
localities, some of the specimens reexamined). Jordan and Dickerson, 1908
(Tampico, Mexico). Hildebrand and Schroeder, 1928 (Chesapeak Bay, many localities).[Gins.
1933b]
Distribution:
From Long Island, NY to Tampico, Mexico[Gin.33]; it is absent from southeastern
Florida and the Keys [B&R 68]
Biology: Dahlberg, M.D. et al 1973 (ecology and reproduction
- Georgia; compared with ginsburgi)
ginsburgi Hildebrand
and Schroeder 1928:324 type USNM 87655 Type locality Chesapeake Bay. [H&S,
1928]
Biology: Dahlberg et al 1973( reproduction and ecology
in Georgia; compared with bosci)
heterolepidotum Hoese
1971:76 (unpublished MS name)Type SIO 67-274 (formerly UA 65-33) Type locality
Punta Peñasco, Sonora, Mexico. Conglomerate reef, midtidal pool. Collected
July 27, 1965.
longipala Ginsburg
1933a:18; holotype USNM 86158; Type locality: Boca Grande, Florida, 2 April
1917; taken by steamer Grampus. [Ginsburg 1933a]
Subgenus Austrogobius
Austrogobius de Buen
1950: 122 (name introduced); 1951: 64-68 Type species: Gobiosoma parri
Ginsburg 1933, by monotypy. [B&R 1968]
parri Ginsburg 1933b:
44-46; Holotype: USNM 93177; Type locality: Pocitos, Uruguay. Fowler 1943:
330 (five specimens from Uruguay; eight dorsal spines in error). Dawson 1963:
585 (in subgenus Dilepidion, name only). [B&R 1968]
Gobius sp.: Barattini
1942: 13 (coast of Uruguay). [B&R 1968]
Austrogobius parri:
de Buen 1950: 122 (synonymy; announcement of genus); 1951; 63-68, pl. 1 (made
type of new genus Austrogobius; redescription, illustrated). [B&R
1968]
Garmannia parri: Robins
1960: 284 (Austrogobius synonymized with Garmannia). Robins
and Bohlke 1964:
5 (retained in Garmannia). [B&R 1968]
grosvenori (Robins)
1964:399-402; holotype ANSP 101166; Paratypes UMML 14380; Type locality: Boca
Chita Pass, Dade Co., FL; Cervigon 1966: 743-745, fig. 313 (three specimens
from Cubagua Island, Venezuela). Caldwell 1966: 70 (specimens from Palisadoes,
Jamaica). [B&R 1968][Robins 1964]
hemigymnum (Eigenmann
and Eigenman) 1888(1889)
Gobius hemigymnus
Eigenmann and Eigenmann 1888(1889): 66-67; holotype MCZ no. 27124; Type locality:
West Indies, exact location unknown). [B&R 1968] [Eigenmann and Eigenmann
1888(1889)]
Garmannia hemigymna:
Ginsburg 1933b: 57-58 (placed in new subgenus Risor). Fowler 1940:
791 (two specimens from Rio de Janeiro). Fowler 1941: 177 (Rio de Janeiro;
in check list). Fowler 1951: 30-31 (redescription on basis of two specimens
from Rio de Janeiro). Robins 1960: 284 ( removed from genus Risor; characters
summarized). [B&R 1968]
Garmannia mediocricula
Ginsburg 1942: 365-366 Type locality: Rio de Janeiro; description, based on
same material as in Fowler's references above). Robins 1964: 402 (compared
with grosvenori). [B&R 1968?] [Ginsburg 1942]
spilotum (Ginsburg)
1939a:62
Garmannia spilota
Ginsburg 1939a: 62; holotype USNM 81828; Type locality: Colon, Panama, in
tidepools. Robins 1960: 284 (compared to Garmannia saucra). Robins
1964: 402 (compared to grosvenori).[Ginsburg 1939a][B&R 1968]
Subgenus Garmannia
Garmannia Jordan and
Evermann, in Jordan et al. 1895: 497 Type species: Gobius paradoxus
Gánther 1861:372,
by original designation.[B&R 1968]
Gerhardinus Meek and
Hildebrand 1928: 889 Type species Gerhardinus nudus Meek and Hildebrand
1928, by original designation.[B&R 1968] [Meek and Hildebrand 1928]
Gobiolepis Ginsburg
1944: 379 Type species: Garmannia hildebrandi Ginsburg 1939a, by original
designation.[B&R 1968][Ginsburg 1944]
Gobiohelpis Ginsburg
1944: 380 Type species: Garmannia spes Ginsburg 1939a:69, by original
designation.[B&R 1968][Ginsburg 1939a][Ginsburg 1944]
Gobiculina Ginsburg
1944: 380 Type species: Garmannia homochroma Ginsburg 1939:62, by original
designation. [B&R 1968][Ginsburg]
chiquita (Jenkins
and Evermann) 1889
Gobius chiquita Jenkins
and Evermann 1889: 146; Holotype USNM 39634; Type locality: Guaymas, Mexico;
spines erroneously listed as six. [B&R 1968]
hildebrandi (Ginsburg
1939a)
Garmannia hildebrandi
Ginsburg 1939a: 61-62; holotype USNM 107297; Type locality: Gatun Locks, upper
lock, Atlantic side of Panama Canal; also from Pedro Miguel Locks, Pacific
side. Hildebrand 1939: 30, 33 (Gatun and Pedro Miguel Locks, report on the
type material). Fowler 1944: 518 (compiled). Ginsburg 1942: 366 (compared
with Garmannia mediocricula). Ginsburg 1944: 379 (made type of subgenus
Gobiolepis; characters). Ginsburg 1945: 136-137 (caudal-ray counts).
Robins 1960: 284 (compared with Garmannia saucra). Robins 1964: 402
(compared with Garmannia grosvenori).[Ginsburg 1939a] [B&R 1968]
homochroma (Ginsburg)
1939a
Garmannia homochroma
Ginsburg 1939a: 62-63; Holotype USNM 107296; Type locality: Pedro Miguel Locks
Pacific slope of Panama. Hildebrand 1939: 33 (listed ten specimens as type
material from Pedro Miguel Locks). [B&R 1968][Ginsburg 1939a]
nudum (Meek and Hildebrand)
1928
Gerhardinus nudus
Meek and Hildebrand 1928:889, pl. 88; holotype USNM 81834; Type locality:
tidepools, Panama City, Pacific Coast of Panama; type of genus Gerhardinus.
Brittan 1966: 132 (Mata Limon estuary, Costa Rica). [B&R 1968] [Meek and
Hildebrand 1928]
Gobiosoma nudum: Ginsburg
1933b: 46-47 (Gerhardinus placed as subgenus of gobiosoma; characters, relationships,
range extended to Peru). Hildebrand 1939: 43 (lower chambers, Miraflores Locks,
Panama). Fowler 1944: 518 (compiled). Rubinoff 1963: 140-145 (distribution
in Panama). [B&R 1968]
paradoxum
(Günther) 1861a
Gobius paradoxus Günther
1861a: 372; Type _____ (not listed in original publication); Type locality:
western coast of Central America; dorsal spines erroneously listed as six;
1861b: 549-550 (description); 1862: 327 (repeat of original description).
Gilbert and Starkes 1904: pl. 38 (good illustration of type species). [B&R
1968] [Gánther 1861a]
robustum Ginsburg
1933a:15-16; Type USNM 92802; Type locality Laguna Madre, near Corpus Pass,
Texas [Gin. 1933]
Biology: Springer & McErlean 1961: 73-75;
Springer & McErlean 1961; Dawson 1966: 384-392; Zavala Garcia 1988[1989]
(early life history & larval distribution, Mexico)
Distribution: Gulf Coast from southern Texas
to Key West, FL and alopng the Atlantic coast to Miami, northward to Cape
Canaveral, FL [B&R 68]
ERRORS: Record from Bahia, Brazil most likely
G. parri[B&R 68]; record from Jamaica by Caldwell 1966:70 is Psilotris
alepis Ginsburg [B&R 68]
schultzi (Ginsburg)
1944
Garmannia schultzi
Ginsburg 1944: 375-378; holotype USNM 121546; paratypes (6) USNM 121547; Type
locality: Venezula, Lake Maracaibo, 7 km S of Maracaibo City. Schultz 1949:
185 (collections from Maracaibo region of Venezula). Robins 1960: 280 (contrasted
with Garmannia saucra).[B&R 1968][Ginsburg 1944]
spes (Ginsburg) 1939a
Garmannia spes 1939a:
62; holotype USNM 107299; Type locality: Canal Zone, dry dock, Mt. Hope. Ginsburg
1942: 366 (contrasted with Garmannia mediocricula). Ginsburg 1944:
377-378, 380 (data on specimens from Venezula; made type species of new subgenus
Gobiohelpis). Fowler 1044: 518 (compiled). Schultz 1949: 185 (collection
from near Sinamaica, Venezula). Caldwell et al. 1959: 28 (recorded from Costa
Rica). Robins 1960: 284 (contrasted with Garmannia saucra). [B&R
1968][Ginsburg 1939a]
Garmannia hildebrandi:
Caldwell 1966: 70 (misidentification, collection from Palisadoes, Jamaica).
[B&R 1968]
yucatanum Dawson 1971:
433-437; holotype USNM 205368; type locality: Mexico, Quintana Roo, south
side of harbor of Cd. Chetumal; 1 meter.
Subgenus Gobiolepis
This subgenus is included because of the new species, which are being described and placed in it. Several other species above also belong here but are being retained in their present subgenera until the manuscripts appear in press.
Gobiolepis Ginsburg 1944
Originally defined as a subgenus
of Garmannia by Ginsburg, Type species Garmannia hildebrandi Ginsburg
1939. Now defined by Hoese '71 (unpublished thesis) as a subgenus of Gobiosoma
and to include the following species: G. hildebrandi, G. birdsongi
(sp. nov.), G. chiquite, G. zacae (sp. nov.), G. grosvenori, G. homochroma.
birdsongi Hoese 1971(unpublished MS name):Holotype ANSP
(formerly UMML 23466) type locality a mud-covered, rocky flat and stream at
base of spillway in Miraflores Locks of the Panama Canal.
zacae Hoese 1971(unpublished MS name): Holotype SU
60374; Type locality - Panama, Bahía Honda.
Subgenus Elacatinus
Elacatinus Jordan
1904: 542; Type species: Gobiosoma oceanops Jordan 1904, by original
designation and by monotypy. This subgenus frequently has been misspelled
in the literature, Elecatinus and Lactinius being the most common
misspellings [B&R 1968]
oceanops Jordan 1904
Elacatinus oceanops
Jordan 1904: 542, pl.2, fig.3 Holotype Stanford Nat. Hist. Mus. 8365, cotype
2757; Type locality: Garden Key, Tortugas, Florida; habitat on coral heads
at depths of 3-8 ft.(original description) [Feddern 1967]
Gobiosoma oceanops: Ginsburg
1933: 2, 19, 21-22. Briggs 1958: 288 (range). Duarte-Bello 1959: 123 (range).[Feddern
1967] Hildebrand, Chavez and Compton 1964: 124 (Alacran Reef, off Yucatan).
[B&R 1968]
Lactinus oceanhops:
Fischer 1955: 36-37, 70 (name misspelled; general and spawning behavior in
aquaria) [Feddern 1967] Straughan 1956: 157 fig. (popular notes on species
in the aquarium) [B&R 1968]
Lactinius oceanops:
Straughan 1959: 140-144, fig. 62 & 63 (name misspelled). Straughan 1964:
165-169, fig. 21, 56, 72, 73, & 75 (spawning in aquarium) [Feddern 1967]
Elecatinus oceanops:
Herold 1961: 235, pl. 100 (name misspelled). Limbaugh 1961: 43 (parasite picking
behavior). [Feddern 1967]
?Elacatinus oceanops:
Wikler 1962: 201-205 (general and spawning behavior; description and fig.
of egg and larve). [Fedder 1967]
Elecatinus oceanhops:
Menten 1962: 246, figs. (misspelled at beginning of paper, notes on spawning
in aquarium). [B&R 1968]
Elacatinus oceanops:
Schroeder 1965: 110-112 (Alligator Reef, Florida Keys; parasite removing habits).
Eibl-Eibesfeldt 1967: fig. 1. Starck and Davis 1966: 340, figs 21 & 22
(inactive at night). [B&R 1968]
not Elacatinus oceanops:
Breder 1927: 83 (West Indies). Fowler 1944: 451 (Bahamas). Eibl-Eibesfeldt
1955: 207-209, figs. 5-7 (description of parasite removing at Bonaire, Dutch
West Indies) [Feddern 1967]
not Gobiosoma oceanops
Erdman 1956: 335 (Jamaica; Haiti; Puerto Rico). [Feddern 1967]
chancei Beebe &
Hollister 1933
Gobiosoma chancei
Beebe and Hollister 1933: 87-88, fig. 17; holotype USNM 170955; type locality:
Grenada, British West Indies, St. George's Bay. [Beebe &Hollister 1933]
Ginsburg 1933b: 25 (synonymized in error with G. horsti). [B&R
1968]
Gobiosoma horsti:
Herre 1942: 14[298] (Hastings, Barbados, taken from a sponge). Cervigon 1966:
740-741 (in part: figure and account based on chancei, reference on
horsti, and Tortugas part of range based on xanthipora). [B&R
1968]
atronasum Böhlke & Robbins 1968
Gobiosoma (Elacatinus)
astronasum Bohlke
and Robins 1968: 88-90, fig. 6; holotype UF 15427; Type locality: Bahama Islands,
at the eastern edge of the Great Bahama Bank. [B&R 1968]
evelynae Böhlke & Robbins 1968
Gobiosoma (Elacatinus) evelynae Böhlke and Robins
1968: 103-109, fig. 10; holotype ANSP 110559; Type locality: sta. 496, Bahamas,
Great Bahama Bank, Nassau vicinity, Green Cay (N of Rose Island), ca. 1/4
mi. NNW of Cay; ca. 50 ft. [B&R 1968]
Elacatinus oceanops:
Eibl-Eibesfeldt 1957: 38 (Bonaire; discussion of cleaning, with drawing of
six gobies cleaning an Epinephelus striatus, including inside of mouth
and under operculum; probably, although not certainly, evelynae). [B&R
1968]
figaro Sazima, Moura and Rosa 1996
Elacatinus figaro
Sazima, Moura and Rosa, 1996:33-38, figs. 1-6; Holotype MZUSP 50859; Type
locality: Pedra do Navio, Iiha Anchieta, Ubatuba, Sao Paulo, Brazil, depth
5 m, 02 May 1996.
genie Böhlke & Robbins 1968
Gobiosoma (Elacatinus)
genie Böhlke & Robins 1968: 109-114, fig. 11; holotype ANSP 110574;
Type locality: sta. 243, Bahamas, Great Bahama Bank, Nassau vicinity, between
North Cay and the small cay just off Long Cay; 6-13 ft. [B&R 1968]
Elacatinus oceanops:
Breder 1927: 83 (Saddle Rock and Washerwoman Cut, Bahamas; probably of this
species). Fowler 1944: 451 (in checklist; based on Breder 1927 ref.). Feder
1966: 342-343, 360 (cleaning habits in the Bahamas; identification uncertain
but probably this species). [B&R 1968]
horsti Metzelaar 1922
Gobiosoma horsti Metzelaar
1922: 139, fig. 2; Lectotype Zool. Mus. Amsterdam 446; Type locality: Caracasbay,
SW part of Curacao, from sponge. Beebe and Tee-Van 1928: 224, fig. 17 (specimens
from Port-au-Prince Bay, Haiti; descriptive information and habitat based
on more than one species). Jordan, Evermann and Clark 1930: 446 (compiled).
Beebe and Hollister 1931: 88 (compared to chancei). Ginsburg 1933b:
22-25 (in part: chancei incorrectly included in synonymy). Fowler 1944:
472 (in part: Grenada record refers to chancei, Tortugas record to
xanthiprora). Fowler 1952: 109 (Haiti; compiled). Briggs 1958: 288
(compiled; in part: Tortugas record based on xanthipora). [B&R
1968]
illecebrosum Böhlke & Robbins 1968
Gobiosoma (Elacatinus)
illecebrosum Böhlke & Robins 1968: 114-116, fig. 12; holotype
ANSP 110693; Type locality: sta. WAS-CARIB-8, Yucatan, Banco Chinchorro, around
Cay Norte; 15-20 ft. [B&R 1968]
louisae Böhlke & Robbins 1968
Gobiosoma (Elecatinus)
lousiae Böhlke and Robins 1968: 90-94, fig.7; holotype ANSP 110691;
Type locality: sta. 14 (G64-43), W side of Grand Cayman Island at dropoff,
offshore from N side of Georgetown; 100-115 ft. [B&R 1968]
prochilos Böhlke & Robbins 1968
Gobiosoma (Elacatinus)
prochilos Böhlke and Robins 1968: 97-100, fig.9; holotype ANSP 110628;
Type locality: sta. TE-57, St. Barthelemy, Port de Gustavia, off the rocky
cliffs just S of Anse Galet; 40-45 ft. [B&R 1968]
randalli Böhlke & Robbins 1968
Gobiosoma (Elacatinus)
randalli Böhlke and Robins 1968: 94-97, fig. 8; holotype ANSP 110672;
Type locality: sta TE-25, Grenadines, St. Vincent Island, N end of Anse Mahaut,
just S of Cumberland Bay; 20-25 ft. [B&R 1968]
tenox Böhlke & Robbins 1968
Gobiosoma (Elacatinus)
tenox Bohlke
and Robins 1968: 83-85, fig. 4; holotype ANSP 110688; Type locality: sta.
TE-42, Dominica, Prince Rupert Bay, Portsmouth; 50ft. [B&R 1968]
xanthiprora Bohlke
& Robbins 1968
Gobiosoma (Elacatinus)
xanthipora Böhlke and Robins 1968: 85-87, fig.5; holotype ANSP 110898;
Type locality: Florida, Monroe Co., 1/4 mi. SE of Alligator Reef Light; about
75 ft. Paratypes: USNM 118105: Tortugas; UMML 8312: Jamaica.
[B&R 1968]
Elacatinus horsti:
Longley and Hildebrand 1941: 227 (in part: synonymy given therein does not
apply to xanthipora, but Tortugas specimens described are that species).
[B&R 1968]
Gobiosoma horsti:
Briggs 1958: 288 (in part: Tortugas record). Bailey et al. 1960: 36
(name only; based on Tortugas record). Caldwell 1966: 70 (specimen from Lime
Cay, Jamaica. [B&R 1968]
Subgenus Tigrigobius
Tigrigobius Fowler
1931: 401 Type species Gobiosoma macrodon Beebe and Tee-Van 1928, by
original designation. [B&R 1968]
Gobicula Ginsburg
1944: 379. Type species: Garmannia gemmata Ginsburg 1939, by original
designation.[Bohlke
& Robbins 1968]
macrodon Beebe &
Tee-Van 1928:226; holotype originally no. 7462 in the Tropical Research Collection
of the New York Zoological Society now at USNM 170896; type locality: Port-au-Prince
Bay, Haiti. [Robins 1958] [Beebe and Tee-Van 1928]
Springer & Woodburn 1960: 72 range
digueti (Pellegrin, 1901)
dilepis (Robins &
Bohlke 1964)
Garmannia (Tigrigogius)
dilepis Robins and Bohlke
1964: 3-6, fig 2; holotype ANSP 100815; Type locality: Sta. 589, coral head
off large bay at NW end of Conception Island, Bahamas, 25 ft.. [B&R 1968][R&B
1964]
gemmatum (Ginsburg
1939a)
Garmannia gemmata Ginsburg
1939a:3-5, fig. 2; holotype USNM 107291; Type locality: Old Providence Island,
Caribbean Sea. Herre 1942: 14 (recorded from Barbados). Ginsburg 1944: 379
(placed in new subgenus Gobicula). Bohlke
and Robins 1960: 92-93 (Gobicula synonymized with subgenus Tigrigobius
of Garmannia; dentition and squamation discussed). Robins 1960:
284 (contrasted with Garmannia saucra). [B&R 1968][Ginsburg 1939]
Garmania gemmata:
Schmitt and Schultz 1940:8 (generic name mispelled; type specimens listed
only).[B&R 1968]
inornatus Bussing
1990:109-110 (type locality: Isla Cocinero, Islas Murcielagos, Costa Rica
(10051'N, 85054'W) depth 9 meters. holotype LACM 32493-43; paratypes also
at LACM.
Gobiosoma brocki (not
Ginsburg) Böhlke and Robins (in part) 1968:67-69 (Isla Gorgona, Columbia;
not including Panama specimens)
Gobiosoma sp., Burgess
& Axelrod, 1984: 2109 (color plate of freshly preserved specimen)
Elacatinus sp. Lopez
& Bussing, 1982.
janssi Bussing 1981:
251-256, fig. 1. (type locality: Bahia Herradura just south of the Gulf of
Nicoya (9038'45"N, 84040'55"W), Pacific coast of Costa Rica) Holotype
LACM 32524-45; paratypes UCR 618-23, UCR 628-11, LACM 32509-37, UCR 638-3
(other paratypes listed in manuscript)
limbaughi Hoese and Reader 2001
Elacatinus limbaughi Hoese
& Reader 2001 More info to follow soon
multifasciatum Steindachner
1876
Gobius lineatus Poey
1861 (a homonym of Gobius lineatus Jenyns, 1842)[Gins. 1933b]
Gobiosoma multifasciatum
Steindachner 1876:Sitzbar. k. k. Ak. Wiss. Wien 74 (abt.1): 231 and 238 (Ichthyol.
Beit. 5: 183 and 190); Type locality: St. Thomas; Barbados and Bartholomew,
Lesser Antilles. [Gins.1933b]
see: Ginsburg 1933b for additional references.
nesiotus Bussing 1990:110-112;
(type locality: East side of Isla del Coco, 300m SW of Cabo Descubierto; depth
20 meters);holotype LACM 44821-3; paratypes also at LACM.
pallens (Ginsburg)
1939a
Garmannia pallens
Ginsburg 1939a:63; holotype USNM 107327; Type locality: Barahona Harbor, Santo
Domingo, 2-4 ft. [Ginsburg 1939a][B&R 1968]
see: Bohlke
& Robins 1960 for summary & literature
puncticulatum Ginsburg 1938
Gobiosoma punticulatum
Ginsburg 1938: 57-58; holotype Stanford University No. 32963; Type locality:
Los Frailes, near Cape San Lucas, Baja California. Fowler 1944: 518 (in check-list
only). [B&R 1968][Ginsburg 1938]
Gobiosoma rubrifrons
Fowler 1944: 282, 383, fig. 237 (type locality: Saboga, Perlas Islands, Gulf
of Panama). [B&R 1968]
saucrum (Robins 1960)
Garmannia saucra Robins
1960: 281-285, 1 fig.; holotype ANSP 92985; Type locality: South Cay, Jamaica,
3 miles south of Port Royal, north side of cay in 3-5 ft. of water. Caldwell
1966: 70 (compiled) [B&R 1968] [Robins 1960]
zebrella (Robins)
1958
Garmannia zebrella Robins 1958:192-194; holotype
ANSP 53387; type locality: Monos Island, Trinidad. [Robins 1958]
Genus Gobulus
Gobulus Ginsburg,
1933a: 12. Type species: Gobulus crescentalis (Gilbert) = Gobiosoma
cresentalis Gilbert, 1892 {see note in bibliography on Gilbert}
crescentalis (Gilbert)
1892:557
Gobiosoma cresentalis
Gilbert, 1892:557 [Gin. 1933a] Type locality: Albatross Exp. Sta. number 2825;
Gulf of California, lat. 24022'15"N., long. 110019'15"W.,
7 fathoms [Gilbert 1892]
Gobiosoma crescentale
Jordan and Everman 1898:2259 U.S. Nat. Mus. Bull. 47, pt. 3 (erroneously said
to have been taken in 79 fathoms) [Gin. 1933a]
Gobiosoma cresentale
Pellegrin 1901:162 Bull. Mus. Hist. Nat. Paris, vol 7, (Gulf of California)
[Gin. 1933a]
Gobiosoma crescentale
Osburn and Nichols 1916:175 Bull. Amer. Mus. Nat. Hist., vol. 35 (Agua Verde
Bay, Gulf of California)[Gin. 1933] [Osburn and Nichols 1916]
birdsongi Hoese and Reader 2001 more info soon
hancocki Ginsburg
1938: 118-119; holotype in USNM; Type locality: Secus Island, Panama; shore,
coral tide flat. [Ginsburg 1938]
myersi Ginsburg 1939a:63;
holotype USNM 107283; type locality: Gulf of Mexico, off Cape Sable; Albatross
station 2374, lat. 29011'30", long. 85029', 26
fathoms. [Ginsburg 1939a] Note: according to Hastings 1983 the type locality
is the extreme northeastern Gulf of Mexico. Ginsburg had the correct latitude
and longitude but listed the wrong location for those marks. [Hastings 1983]
Genus Gymneleotris
Gymneleotris Bleeker
1874: 304 Type species: Eleotris seminudus Gánther 1864, by original
designation. [B&R 1968]
seminudus (Gánther 1864)
Eleotris seminudus Gá