signatus Poey 1876   

brevispinis Ginsburg 1939  

carri Fowler 1945

crocatus Birdsong 1968

curtus Ginsburg 1939

cyclolepis Gilbert 1891

emblematicus (Jordan and Gilbert) 1882   

erectus Ginsburg 1938

gulosus (Girard) 1858

meeki Evermann and Marsh 1900

microlepis Longley and Hildebrand 1940

miraflorensis Gilbert and Starks 1904

tabogensis Meek and Hildebrand 1928

thalassinus (Jordan and Gilbert 1882a)